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Dung beetle (Scarabaeidae) pushing ball of dung on Venetia Limpopo Reserve, Limpopo Province, South Africa
The Pinnacles, limestone formations. Nambung National Park, near Cervantes, Western Australia, November 2015
Mount Lidgbird (777 m) and Kentia palms (Howea forsteriana) with two tall Curly palm trees (Howea belmoreana) Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site
Polar bear (Ursus maritimus) silhouetted against setting sun, Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
North American Bobcat (Lynx rufus) stalking along Madison River. Yellowstone National Park, Wyoming, USA. January
Cinnamon bear, subspecies of black bear (Ursus americanus cinnamomum) cub climbing tree, Yellowstone National Park, Wyoming, USA, May
RF- Mawddach Estuary at low tide, Barmouth, Snowdonia National Park, Gwynedd, Wales, May 2012
RF- River Plym flowing through Dewerstone Wood, Dartmoor National Park, Devon, England, UK, October
Chilean flamingos (Phoenicopterus chilensis) group of three in flight, Lago Amarga, Torres del Paine National Park, Patagonia, Chile
Red squirrel (Sciurus vulgaris) on old pine tree, looking through autumn birch foliage, Scotland, UK, October
Red Squirrel (Sciurus vulgaris) reaching up and standing on hind legs. Cairngorms National Park, Highlands, Scotland, UK, March
Red squirrel (Sciurus vulgaris) in summer coat on Scots pine tree trunk, Highlands, Scotland
Capercailie (Tetrao urogallus) male displaying from branch of Scots pine tree. Cairngorms National Park, Highlands, Scotland, UK, March
Sessile Oak (Quercus petraea) fallen oak leaf and acorns on moss, Highlands, Scotland, October
Young ground squirrels (Xerus inauris) playing, Kgalagadi Transfrontier Park, South Africa. Non-ex
Bromeliads (Bromeliaceae) in flower in rainforest, Salto Morato Nature Reserve / RPPN Salto Morato, Guaraquecaba, Parana, Brazil
Lilac-breasted roller (Coracias caudata) stretching wings on acacia branch, Masai Mara Game Reserve, Kenya. November
Yellow mongoose (Cynictis penicillata) standing on hind legs with young, Kgalagadi Transfrontier Park, South Africa, February
Meerkats (Suricata suricatta) standing alert, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Chamois (Rupicapra rupicapra) running down ridge, in early morning light. Lausons Valley, Gran Paradiso National Park, Italy, September
Helmcken Falls, Wells Gray Provincial Park, British Columbia, Canada, July
River Gave de Gavarnie with Cirque du Gavarnie in the distance, Pyrenees National Park, France, June 2013
European common frog (Rana temporaria) on rock by waterfall, near Lac des Gloriettes dam, Pyrenees National Park, France, June
Black veined white butterfly (Aporia crataegi) on grass, Gavarnie-Gedre, Pyrenees National Park, France, June
Male Ostrich (Struthio camelus) protecting chicks from the sun with its wings, Samburu National Reserve, Kenya, Africa
Female Grizzly bear (Ursus arctos horribilis) standing up, with two cubs nearby, Khutzeymateen Grizzly Bear Sanctuary, British Columbia, Canada, June
Lion (Panthera leo) male shaking his head after rain, Masai-Mara Game Reserve, Kenya
Bighorn sheep (Ovis canadensis) close up rams head shot, Yellowstone National Park, USA, February
Topi (Damaliscus korrigum) at sunset, Masai-Mara Game Reserve, Kenya
Indri (Indri indri) calling, Andasibe-Mantadia NP, Madagascar
Fringed leaf frog (Cruziohyla craspedopus) on branch, Yasuni National Park, Orellana, Ecuador
Amazon wood lizard / Guichenots dwarf iguana (Enyalioides laticeps) portrait, Yasuni National Park, Orellana, Ecuador
Brush footed butterfly (Lycorea sp. ) caterpillar with parasitic wasp cocoons on silk threadsBrush footed butterfly (Lycorea sp.) caterpillar with parasitic wasp cocoons on silk threads, Yasuni National Park, Orellana, Ecuador
Tiger leg monkey frog / Tiger striped monkey frog (Phyllomedusa tomopterna) portrait, on stem, Yasuni National Park, Orellana, Ecuador
Giraffe weevil (Trachelophorus giraffa) Andasibe-Mantadia National Park, Moramanga, Madagascar
Indian peafowl (Pavo cristatus) peacock walking and displaying feathers, Yala National Park, Southern Province, Sri Lanka
Fire salamander (Salamandra salamandra) female almost ready to give birth to her larvae into the stream, Apennines. Antola Regional Park, Italy
Rangimairewhenua or Blue Lake, Nelson Lakes National Park, Southern Alps, New Zealand, February 2013. Photographed for The Freshwater Project
Dogbane beetle (Chrysochus auratus) on dogbane, Crossways Preserve, Philadelphia, Pennsylvania, USA, August
Passionflower (Passiflora cubalensis) Yanacocha Reserve, Ecuador
Elephant Hawkmoth (Deilephila elpenor) close up of head showing proboscis. Note the pollen grains attached to its hairs. Peak District National Park, Derbyshire, UK. August. Focus Stacked
Velvet shank / Winter fungus (Flammulina velutipes), growing on dead tree. Peak District National Park, UK. December. Focus stacked image
Small copper butterfly (Lycaena phlaeas) in meadow. Peak District National Park, Derbyshire, UK. May
Porcelain fungus (Oudemansiella mucida) growing on dead Beech tree (Fagus sylvatica). Peak District National Park, Derbyshire, UK. October
Small tortoiseshell butterfly (Aglais urticae) feeding on Marjoram (Origanum vulgare). Peak District National Park, Derbyshire, UK. July
Ash Tree (Fraxinus excelsior) growing in a field. Peak District National Park, Derbyshire, UK. August
Mountain hare (Lepus timidus) grooming itself, with back foot raised, Cairngorms National Park, Scotland, UK, February. Highly Commended in the Animal Behaviour category of the British Wildlife
Red deer (Cervus elaphus) stag in autumn woodlands, Bradgate Park, Leicestershire, UK, October