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Magnificent hummingbird (Eugenes fulgens) female, Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Fiery throated hummingbird (Panterpe insignis) an endemic bird species. Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Massive old growth oak trees of the cloud forest, Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica. Small repro only
Mountain landscape of Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Fossil leaves of seed ferns, a kind of extinct plant, Joggins Fossil Cliffs UNESCO World Heritage Site, Bay of Fundy, Nova Scotia, Canada. May 2017
Semipalmated sandpiper (Calidris pusilla) flock in flight over water, Johnsons Mills Reserve, Bay of Fundy, New Brunswick, Canada, July
Alpine ibex (Capra ibex) close-up portrait. Gran Paradiso National Park, the Alps, Italy. January. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Cliff parakeets (Myiopsitta luchsi) perched on cactus, Red-fronted Macaw Community Nature Reserve, Omerque, Bolivia
RF - Mute swan (Cygnus olor) taking off from a pond Valkenhorst Nature Reserve, Valkenswaard, the Netherlands June
Kiang (Equus kiang) walking, Sanjiangyuan National Nature Reserve, Qinghai Hoh Xil UNESCO World Heritage Site, Qinghai-Tibet Plateau, Qinghai Province, China
Blue poppy (Meconopsis horridula) Sanjiangyuan National Nature Reserve, Qinghai Hoh Xil UNESCO World Heritage Site, Qinghai-Tibet Plateau, Qinghai Province, China
Coral Rabbitfish (Siganus corallinus) portrait, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Crowned dwarf caiman (Paleosuchus trigonatus) male resting on a log. Yasuni National Park, Orellana, Ecuador
RF- Cheetah (Acinonyx jubatus) cubs aged 6 months, Masai-Mara Game Reserve, Kenya. Vulnerable species
Land crab (Geosesarma sp. ) which raids Pitcher plant (Nepenthes ampullaria) for preyLand crab (Geosesarma sp.) which raids Pitcher plant (Nepenthes ampullaria) for prey, Sarawak, Borneo
Wild horses, Monument valley tribal park, Navajo reserve, Utah, USA. April
Polar bear (Ursus maritimus), female with two cubs, on a barrier island outside Kaktovik, Alaska, USA
Przewalski horse (Equus ferus przewalski) Khustain Nuruu National Park, Mongolia. December
Amur leopard (Panthera pardus orientalis) Land of the Leopard National Park, Primorsky Krai, Far East Russia. March
Green huntsman spider (Olios sp), Intervales State Park, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Portrait of a Golden snub-nosed monkey 1+Rhinopithecus roxellana+2 full frame of the face, Foping Nature Reserve, Shaanxi, China. Endangered species
Golden snub-nosed monkey 1+Rhinopithecus roxellana+2 sitting in tree, Foping Nature Reserve, Shaanxi, China. Endangered species
Golden snub-nosed monkey (Rhinopithecus roxellana) jumping from branch to branch, Foping Nature Reserve, Shaanxi, China. Endangered species
Chimpanzee (Pan troglodytes schweinfurthii) male, scratching its leg, National Park, Uganda
RF - Mountain gorilla (Gorilla beringei beringei) silverback male, portrait, member of the Nyakagezi group, Mgahinga National Park, Uganda. Critically endangered
Mountain gorilla (Gorilla beringei beringei) silverback male, portrait, member of the Nyakagezi group, Mgahinga National Park, Uganda. Critically endangered
Yunnan snub-nosed monkeys (Rhinopithecus bieti) adult and young, Ta Cheng Nature reserve, Yunnan, China. October
Golden pheasant (Chrysolophus pictus) displaying to female pheasants at Yangxian Nature Reserve, Shaanxi, China, September
Red jungle fowl (Gallus gallus) Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China, April
Bluespine unicornfish (Naso unicornis) Shark Reef to Jolande, Ras Mohammed National Park, Egypt, Red Sea
Indian giant squirrel (Ratufa indica) Kaziranga National Park, Assam, India
Pheasant (Phasianus colchicus) male calling, East Frisian Islands, Wittbulten National Park. Wadden Sea UNESCO World Heritage Site, Germany, June
Spotted hyena (Crocuta crocuta), cubs together by den, Masai-Mara Game Reserve, Kenya
Hippopotamus (Hippopotamus amphibius), female with calf on land, Masai-Mara Game Reserve, Kenya
Lion (Panthera leo), females hunting buffalo, Masai-Mara Game Reserve, Kenya
Crab-eating fox (Cerdocyon thous) Kaa-Lya National Park, South East Bolivia
Conovers tuco-tuco (Ctenomys conoveri) digging burrow Kaa-Iya National Park, Bolivia
Brown bearded sugar ant (Camponotus terebrans) William Bay National Park, Western Australia. Meetyourneighbours.net project
Reindeer (Rangifer tarandus) female from reintroduced herd on Cairngorm Mountains, Cairngorm National Park, Speyside, Scotland, December
European otter (Lutra lutra) with snow ball, Bavarian Forest National Park, Germany, January. Captive
Waterviolet / featherfoil (Hottonia palustris) at sunset, Groot Schietveld, Wuustwezel, Belgium
Red-fronted macaw (Ara rubrogenys) two in flight, Red-fronted Macaw Community Nature Reserve, Omerque, Bolivia
Black spider monkey (Ateles chamek) in tree, Madidi National Park, Bolivia
BlackbuckA(Antilope cervicapra) herd with males and females, Velavadar national parkBlackbuck(Antilope cervicapra) herd with males and females, Velavadar national park, Gujarat, India
Tiger (Panthera tigris), female with large cubs, Ranthambhore National Park, Rajasthan, India
Tiger (Panthera tigris), rolling on back and relaxing after feeding, Ranthambhore National Park, Rajasthan, India
Nilgai (Boselaphus tragocamelus), male walking through water, Keoladeo Ghana National Park, Bharatpur, Rajasthan, India
Grey heron (Ardea cinerea) squabbling with Little cormorant (Microcarbo / Phalacrocorax niger) Keoladeo national park, Bharatpur, Rajasthan, India