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Spiny / Yellow Seahorse (Hippocampus guttulatus). Studland Bay, Dorset, UK, September
Sunfish (Mola mola) basking at the surface on summers day. Penzance, Cornwall, England, British Isles, August. North East Atlantic Ocean
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
Sohal surgeonfish (Acanthurus sohal), Red Sea, Egypt. January
Rock flagtail (Kuhlia rupestris) fish in river on the Southern Lagoon, Forgotten Coast, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos
Rainbow trout (Oncorhynchus mykiss) in green turbid waters, Gunnison River, Colorado, USA, April
Piraputanga, (Brycon hilarii) reflected on the water surface, Aquario Natural, Bonito, Mato Grosso do Sul, Brazil
Bohar snapper (Lutjanus bohar) school at Ras Mohammed, Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea
Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Yellow warty frogfish (Antennarius maculatus) Seraya, Tulamben area, Bali, Indonesia. Java Sea
Silver Drummer (Kyphosus sydneyanus) fish off the coast of Neds Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Giant Manta Ray (Manta birostris), with Remoras (Echeneididae) Socorro Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March
Young Stone sculpin (Paracottus knerii), on endemic sponge (Lubomirskia baicalensis), Lake Baikal, Siberia, Russia
Tropical Flounder (Bothus mancus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
European Chubb (Squalius / Leuciscus cephalus) in cave, Plitvice Lakes National Park, Croatia. November
Orange anemonefish (Amphiprion sandaracinos) in its host Mertens carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea
Flame anthias (Pseudanthias ignitus) displaying to the more drab female (above) during courtship in the evening. Similan Islands, Thailand. Andaman Sea, Indian Ocean
Great hammerhead shark (Sphyrna mokarran) female at night with Bar jacks (Caranx ruber), South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Patagonian redfish (Sebastes oculatus) about 4cm, Comau Fjord, Patagonia, Chile
Mandarinfish (Synchiropus splendidus) Yap, Micronesia
Giant moray eel (Gymnothorax javanicus) opening its mouth to breathe, at night on the barge wreck, Gubal Island, northern Red Sea
Dwarf lionfish (Dendrochirus brachypterus) opens its awingsa (in fact enlarged pectoralDwarf lionfish (Dendrochirus brachypterus) opens its wings (in fact enlarged pectoral fins) Seraya, Tulamben area, Bali, Indonesia, Java Sea
Bignose unicornfish (Naso vlamingii) profile of male displaying (the blue stripe at the bottom of the frame is the tail of another individual) Baa Atoll, Maldives. Indian Ocean
Blackspotted stingray (Taeniurops meyeni) with a remora (Echeneis naucrates) attached to its belly swimming in the open water with a diver in the background. Ari Atoll, Maldives, Indian Ocean
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Galapagos barnacle blenny (Acanthemblemaria castroi) looking out from its home in an old barnacle shell. Cape Marshall, Isabela Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Clarks anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia
Bigeye trevally (Caranx sexfasciatus) forming a circular shoal in blue water, on the drop off of a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean
Black Bream (Spondyliosoma cantharus) washed up on shore, Sark, British Channel Islands, August
Short snouted seahorse juvenile (Hippocampus hippocampus) Sark, British Channel Islands, August
Ling (Molva molva) St Abbs Voluntary Marine Reserve, Scotland (North Sea)
Eggs of Long-spined sea scorpion (Taurulus bubalis) on sea shore, possibly being predated by Painted Topshell, Guernsey, British Channel Islands
John dory (Zeus faber) profile portrait, Channel Islands, UK October
Leopard-spotted Goby (Thorogobius ephippiatus). Channel Islands, UK, June
Cuckoo wrasse {Labrus mixtus / bicamulatus} male, Channel Islands, UK
RF - Orange skunk clownfish (Amphiprion sandaracinos) in anemone, Puerto Galera, Philippines
Big-eye or Goggle-eye (Priacanthus hamrur) and Crown squirrelfish (Sargocentron diadema) on wreck colonised by corals, Red Sea, Egypt. January
Kaleidoscopic image of coral reef with soft corals (Dendropnethya) shoal of Yellow back fusilliers (Caesio teres) West Papua, Indonesia
RF - Coral reef with Soft corals (Dendronephthya sp) and fish, West Papua, Indonesia
Spinecheek anemonefish (Premnas biaculeatus) male with bleached anemone, Bunaken National Park, North Sulawesi, Indonesia
Big-eye fish (Priacanthus hamrur). Egypt, Red Sea
Emperor angelfish (Pomacanthus imperator) swimming past coral reef. Egypt, Red Sea
Yellowbar angelfish (Pomacanthus maculosus) Egypt, Red Sea
Coral hind (Cephalopholis miniata) lying in ambush under table coral on reef. Andaman Sea, Thailand
Coral reef scenery with Gorgonian and shoal of Pygmy / Glassy sweepers (Parapriacanthus guentheri) Egypt, Red Sea