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Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Portrait of Blacktip grouper (Epinephelus fasciatus) resting in Barrel sponge (Xestospongia testudinaria) Mabul Island, Sabah, Borneo, Malaysia. Sulawesi Sea
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by a group of Pilotfish (Naucrates ductor). Rocky Island, Egypt. Red Sea
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
Large male Napoleon wrasse (Cheilinus undulatus) yawning. Ras Mohammed, Sinai, Egypt. Red Sea
School of Longnose parrotfish (Hipposcarus harid) swarming over reef. Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
RF- School of Blackfin barracuda (Sphyraena qenie) forming circle in open water at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Black rockfish (Sebastes melanops) in bull kelp (Nereocystis luetkeana), Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada
Klunzingers wrasse (Thalassoma klunzingeri) swimming over the edge of fringing reef where the water depth drops instantly from few feet to hundreds of meters. Ras Mohammed Marine Park, Sinai, Egypt
Male Yellow-headed jawfish (Opistognathus aurifrons) blows out the clutch of eggs he is incubating in his mouth to oxygenate them. East End, Grand Cayman, Cayman Islands, British West Indies
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
RF - Close up detail of the eye and skin pattern of a Giant hawkfish (Cirrhitus rivulatus). Socorro Island, Pacific Ocean
Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean
A Leafy Seadragon (Phycodurus eques), photographed from below. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia. Australia
RF - Banggai cardinalfish (Pterapogon kauderni). Lembeh Strait, North Sulawesi, Indonesia
Citron (lemon coral) goby (Gobiodon citrinus) Red Sea
Two Common butterfly fish (Chaetodon chrysurus) Red Sea, Egypt
Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
Blue and gold fusilier / Scissor tailed fusilier (Caesio caerulaurea) school swimming through shafts of sunlight, West Papua, Indonesia
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Blue triggerfish (Pseudobalistes fuscus) Egypt, Red Sea
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Ruffe (Gymnocephalus cernua) shoal, Danube Delta, Romania, June
Philippines chromis (Chromis scotochiloptera) swimming in front of reef wall with sponges and featherstars, Komodo National Park, Indonesia
Sargassum fish (Histrio histrio) near the surface on floating Sargassum seaweed, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs
Blue spotted trevally (Caranx bucculentus) hunting small fish in the reef, South Ari Atoll, Maldives, Indian Ocean
Red Sea anemonefish (Amphiprion bicinctus) in their home, a Sea anemone (Heteractis magnifica), on a coral reef. Big Brother island, Red Sea. Egypt
Johnsons abyssal seadevil, (Melanocetus johnsonii) deep sea species from Atlantic Ocean off Cape Verde. Captive
Common fangtooth (Anoplogaster cornuta) portrait, deep sea species from Atlantic Ocean off Cape Verde. Captive
Ribbon sawtail fish (Idiacanthus fasciola) from Atlantic Ocean, at a depth of 800-1000m
Dragonfish (Melanostomias melanops) - deep sea specimen from 2000m depth, nr Cape Verde Islands, North Atlantic
Viperfish (Chauliodus sloani) - deep sea specimen from Portugal
Lanternfish (Lepidophanes guentheri) - deepsea species showing bioluminescence
Black Dragonfish (Malacosteus niger)
Deep sea fish (Winteria telescopa) with forward pointing tubular eyes
Angler fish (Melanocetus murrayi) Mid-Atlantic Ridge, North Atlantic Ocean
Deepsea fish {Paraliparis sp. ) North East Atlantic Ocean off Cape Verde Islands depthDeepsea fish {Paraliparis sp.) North East Atlantic Ocean off Cape Verde Islands depth 700-1000m
Bathypelagic Lizard fish (Bathysaurus ferox), deep sea Atlantic ocean
Deep sea Anglerfish female with lure {Himantolophus sp} Atlantic ocean
Deep sea Anglerfish {Melanocoetus sp} female with lure projecting from head to attract prey. Atlantic ocean
Deep sea Anglerfish {Himantolophus sp} female with lure projecting from head to attract prey, Atlantic ocean. Anglerfish have small poorly developed eyes but detect movement
Atlantic Puffin (Fratercula arctica) in flight, coming into land with beak full of sand eels (Ammodytes tobianus). Fair Isle, Shetland Islands, Scotland, UK, July
Three False clownfish (Amphiprion ocellaris) in Sea anemone (Heteractis magnifica) Lighthouse Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean