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RF - Shoal of Slender pinjalo snappers (Pinjalo lewisi) swimming over soft corals on coral reef. West Papua, Indonesia
Ring-tailed Cardinalfish (Apogon aureus) West Papua, Indonesia
Crown spotty blenny (Laiphognathus longispinis) Pak Lap Tsai, Sai Kung East Country Park, Hong Kong, China
Blue shark (Prionace glauca) with Pilot fish (Naucrates ductor) off Halifax, Nova Scotia, Canada. July
Piraputanga (Brycon hilarii) and small Tetra fish (Hyphessobrycon sp. ) Rio Olhio d AguaPiraputanga (Brycon hilarii) and small Tetra fish (Hyphessobrycon sp.) Rio Olhio d Agua, tributary of Rio da Prata, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul
Atlantic salmon (Salmo salar) jumping up waterfall during spawning migration upstream, Ume River, Sweden. July. Photographed for The Freshwater Project
Sargassum fish (Histrio histrio) at the sea surface with floating sargassum weed. Hawaii
Painted cardinalfish (Archamia fucata) Yap, Micronesia
European barracuda (Sphyraena sphyraena) and Bluefish (Pomatomus saltatrix) circling baitball of Atlantic horse mackerel (Trachurus trachurus) Formigas Islets, Azores
Bullethead parrotfish (Chlorurus sordidus) male, close up of eye, whilst sleeping at night. The Alternatives, Sinai, Egypt. Red Sea
Whitebelly damselfish (Amblyglyphidodon leucogaster) pair spawning / laying eggs on a seafan (Annella mollis). Ras Mohammed National Park, Sinai, Egypt. Red Sea
Goby (Bryaninops sp) on coral branch, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Reef and shoal of Jewel fairy basslet (Pseudanthias squamipinnis) South Point dive site, Sanganeb reef, Sudan, Red Sea
Underwater view of African tigerfish (Hydrocynus vittatus), predatory fish in the Okavango river with Water lily (Nymphaea nouchali var. caerulea) and other aquatic plants
Underwater view of Brown bear (Ursus arctos) paw fishing for Sockeye salmon (Oncorhynchus nerka) with paw outstretched. Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Far East Russia. August
Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp)Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp.) diving towards coral reef to avoid predators. North Ari Atoll, Maldives. Indian Ocean
Oriental sweetlips (Plectorhinchus vittatus) pair. South Ari Atoll, Madlives. Indian Ocean
Reticulate stargazer (Uranoscopus sp. ) portrait. Ambon Bay, Ambon, Maluku Archipelago, IndonesiaReticulate stargazer (Uranoscopus sp.) portrait. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Spotfin Burrfish (Chilomycterus reticulatus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Pearlfish (Carapus acus) outside of host Sea cucumber (Stichopus regalis) at night. These species have a commensal relationship, with the fish living in the gut of the sea cucumber. Catalonia, Spain
Common fangtooth (Anoplogaster cornuta) deep sea species from Atlantic Ocean close to Cape Verde. Captive
Spaghetti garden eels (Gorgasia maculata) stretching up out of their burrows on a rubble slope, with Mysiid shrimp swimming around their bodies near the seabed. Dumaguete, Dauin, Negros, Philippines
Bluestreak cleaner wrasse (Labroides dimidiatus) just swimming out of mouth of Oblique / Diagonal banded sweetlips (Plectorhinchus lineatus) whilst cleaning / feeding, New Caledonia. Pacific Ocean
Tassled scorpionfish (Scorpaenopsis oxycephala) swimming just above sea bed, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Lanternfish (Lepidophanes guentheri) - deepsea species showing bioluminescence
A male Leafy Seadragon (Phycodurus eques) swimming. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
Mudskippers {Periophthalmus sp} forage on the mudflats of a mangrove channel, Sundarban Forest, Khulna Province, Bangladesh
Atlantic wolffish {Anarrhichus lupus} Sognefjord, Norway
Shoal of Lake Ohrid bleak (Alburnus scoranza) swimming amongst Giant reeds (Arundo donax) along the shore of Lake Ohrid, Lagadin region, Galicica National Park, Macedonia, June 2009
Ornate wrasse (Thalassoma pavo) shoal, Deserta Grande, Desertas Islands, Madeira, Portugal, August 2009
Wide-eyed flounder (Bothus podas) on seabed, Marine Reserve, Monaco, Mediterranean Sea, July 2009
Scorpionfish (Scorpaena porcus) lying on the artificial reef, Larvotto Marine Reserve, Monaco, Mediterranean Sea, July 2009
Shoal of Mediterranean / Blue damselfish (Chromis chromis) Larvotto Marine Reserve, Monaco, Mediterranean Sea, July 2009
Atlantic bluefin tuna (Thunnus thynnus) portrait, captive, Malta, Mediteranean, May 2009
Atlantic bluefin tuna (Thunnus thynnus) shoal, captive, Malta, Mediteranean, May 2009. WWE INDOOR EXHIBITION
Brown trout (Salmo trutta) at water surface in shallow water migrating upstream, Bornholm, Denmark, November 2008
Close up of Ratfish / Ghost shark (Chimaera monstrosa) Trondheimsfjorden, Norway, February 2009
Dab (Limanda limanda) portrait, Lofoten, Norway, November 2008. WWE BOOK. WWE OUTDOOR EXHIBITION
Two Atlantic wolffish (Anarhichas lupus) looking out of hole, Saltstraumen, Bod, Norway, October 2008
Two Scorpionfish (Scorpaena porcus) lying on artificial reef, Larvotto Marine Reserve, Monaco, Mediterranean Sea, July 2009 WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS
Tench (Tinca tinca) in peat pond, Fribourg, Switzerland, May 2009
Male Zander / Pikeperch (Stizostedion / Sander lucioperca) defending nest, with Chub (Leuciscus cephalus) in the distance, calm part of the Rhine, Schaffhausen, Switzerland, May 2009
European grayling (Thymallus thymallus) pair spawning, the dark one is the male, Lake Thun, Thun, Switzerland, March 2009
European chub (Squalius / Leuciscus cephalus) and Bream (Abramis brama) shoal in calm part of the Rhine, Schaffhausen, Switzerland, February 2009
Atlantic salmon (Salmo salar) migrating upstream to spawn, Umelven, Sweden, July 2009
Anglerfish (Lophius piscatorius) portrait, Saltstraumen, Bod, Norway, October 2008. WWE INDOOR EXHIBITION
Shrimp (Lebbeus polaris) eyeballing an Atlantic wolffish (Anarhichas lupus) Saltstraumen, Bod, Norway, October 2008
Anglerfish (Lophius piscatorius) underside, Saltstraumen, Bod, Norway, October 2008