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Wolf-eel (Anarrhichthys ocellatus) Vancouver Island, British Columbia, Pacific Ocean
Killer whale (Orcinus orca) diving and hunting for herring fish (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii
Dover sole (Solea solea) camouflaged on sea floor, Channel Islands, UK August
Gorgonian / Sea whip coral (Ellisella ceratophyta) with Ring tailed cardinalfish (Apogon aureus) swimming in front, West Papua, Indonesia
Coral reef with soft corals (Dendronephthya sp. ), encrusting sponges and Pygmy sweepersCoral reef with soft corals (Dendronephthya sp.), encrusting sponges and Pygmy sweepers (Parapriacanthus ransonetti) Similan Islands, Andaman Sea, Thailand
Large shoal of Horse mackerel (Megalaspis cordyla) schooling. Rinca, Komodo National Park, Indonesia
Thread-tail dartgoby (Ptereleotris hanae) Sharp Island or Kiu Tsui Chau, Kiu Tsui Country Park, Hong Kong, China
School of Perch (Perca fluviatilis) amongst Water lilies (Nuphar lutea) lit by the suns rays, Lake Bourget, Alps, Savoie, France, June
Paddle-flap scorpionfish (Rhinopias eschmeyeri) Puerto Galera, Philippines
Devil scorpionfish (Scorpaenopsis diabolus) close up of eye, Pak Lap Tsai, Sai Kung, Hong Kong, China
Stellar rockskipper (Entomacrodus stellifer) hiding in a barnacle. Sai Kung East Country Park, Hong Kong, China
Ocean sunfish (Mola mola) off Halifax, Nova Scotia, Canada. July
Atlantic wolfish (Anarhichas lupus) looking out hole, Bay of Fundy, New Brunswick, Canada, July
Atlantic salmon (Salmo salar) portrait, migrating to spawn in river, Gaspe Peninsula, Quebec, Canada, October
Three spot damselfish (Dascyllus trimaculatus) with Sea anemone home, Yap, Micronesia
Common anemonefish (Amphiprion perideraion) with eggs in Magnificent sea anemone (Heteractis magnifica) Yap, Micronesia
Striped mackerel (Rastrelliger kanagurta) with mouths wide open as they swim through the water, filtering zoopankton with their gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) sticking its head into the mouth of a Red Sea fusilier (Caesio lunaris) during a cleaning session. Small Crack, Sha ab Mahmud, Sinai, Egypt. Red Sea
Coral Rabbitfish (Siganus corallinus) portrait, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Cichlids (Cichlidae) in Lake Malawi, Malawi, November. Photographed for The Freshwater Project
Striped mackerel (Rastrelliger kanagurta) shoal swimming with open mouths, filtering for zooplankton. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Bluespine unicornfish (Naso unicornis) Shark Reef to Jolande, Ras Mohammed National Park, Egypt, Red Sea
Trumpetfish (Aulostomus maculatus) using a Spanish Hogfish (Bodianus rufus) as a stalking horse. Spanish Hogfish is eating an arm of a starfish. Bonaire, Leeward Antilles, Caribbean
Pearlfish (Carapus acus) in anus of Sea cucumber (Stichopus regalis). These species have a commensal relationship, with the fish living in the gut of the sea cucumber. Catalonia, Spain
Fangtooth moray eel (Enchelycore anatina) with mouth open, Grand Canaria, Canary Islands, Spain. East Atlantic Ocean
Illuminated netdevil (Linophryne arborifera), female specimen, from deepsea Atlantic Ocean near Cape Verde at a depth of 1900m
Binocular fish {Winteria telescopa}
Female Angler fish {Himantolophus sp} deep sea species
A male Leafy Seadragon (Phycodurus eques) carries a new batch of eggs on his tail. Wool Bay Jetty, Edithburgh, Yorke Peninsula, South Australia, November
Ratfish / Ghost shark (Chimaera monstrosa) on fjord bed, Trondheimsfjorden, Norway, November 2006
Female Atlantic salmon (Salmo salar) River Orkla, Norway, October 2008
Atlantic bluefin tuna (Thunnus thynnus) portrait, captive, Malta, Mediteranean, May 2009 WWE BOOK
Monkfish / Allmouth (Lophius piscotorius) close-up of face, Lofoten, Norway, November 2008
Anglerfish (Lophius piscatorius) Saltstraumen, Bod, Norway, October 2008
Catch of Atlantic mackerel (Scomber scombrus) in fish separator on board Shetland pelagic trawler Charisma, Shetland Isles, Scotland, UK, October 2012
Fraserburgh pelagic trawler Forever Grateful fishing for Atlantic mackerel (Scomber scombrus) close to Papa Stour, Shetland Isles, Scotland, UK, October 2012
Male Black faced blenny (Tripterygion delaisi), part of a sequence showing colour change, Swanage, Dorset, England, UK, May. 2/4
Shanny (Lipophrys pholis) in a crevice in a harbour wall, Bovisand Harbour, Plymouth, Devon, England, UK, July
John dory (Zeus faber), Porthkerris Cove, Cornwall, England, UK, June
Tompot blenny (Parablennius gattorugine) beneath Swanage Pier, Dorset, UK. Photographed during May. English Channel
Brown trout (Salmo trutta) in turbulent water at a weir, River Ettick, Selkirkshire, Scotland, UK, October
Atlantic wolf fish (Anarhichas lupus) in a rock crevice, St Abbs (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, UK, August 2011
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea in the late afternoon, close to Little Brother Island
Snorkeller with shoal of Piraputanga (Brycon hilarii), Rio Olha d Agua at Cabeceira do Prata, Bonito, Mato Grosso do Sul, Brazil
Great Fen / Raft spider (Dolomedes plantarius), adult female eating an invasive species of fish, Western mosquitofish, (Gambusia affinis), Alessandria, Italy, August