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Turnstones (Arenaria interpres) and Purple sandpipers (Calidris maritima) over wintering on the beaches of the Orkney Isles, Scotland, UK, April
Aerial view of nesting Gannets (Morus bassanus) and swirling seas, Hermaness, Shetland, Scotland, UK, October
Dead mans fingers (Alcyonium digitatum) on a rocky reef with view of the sky, Shetland, Scotland, UK, August
Maidens salp (Thetys vagina), west coast of Scotland, UK, July
Aerial view Fin whales (Balaenoptera physalus) lunge-feeding in the southern Sea of Cortez (Gulf of California), Baja California, Mexico
Orca (Orcinus orca) surfacing, Shetland, Scotland, UK, August
Gannet (Morus bassanus) diving for fish, Noss, Shetland Islands, Scotland, UK, August
Autumn ladies tresses (Spiranthes spiralis) Sark, British Channel Islands, August
Lesser spotted catshark / Dogfish shark (Scyliorhinus canicula) amongst seaweed on a maerl bed, South Arran Marine Protected Area, Isle of Arran, Scotland, UK, August
Grey seal (Halichoerus grypus) playful young seal above a kelp forest. Lundy Island, Devon, UK Bristol Channel, August
Grey seal (Halichoerus grypus) young male swimming above Kelp / Oarweed (Laminaria hyperborea) Farne Islands, Northumberland, UK, North Sea, August
Blue shark (Prionace glauca) near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August. Original version of 1439257
California sealion (Zalophus californianus) swimming. Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
Sea otter (Enhydra lutris) floating on its back at the surface among the kelp, Alaska, USA, Gulf of Alaska. Pacific ocean
Galapagos green turtle (Chelonia mydas agassizi) swimming in coastal waters of San Cristobal Island, Galapagos
Sperm whale (Physeter macrocephalus) aerial view. Baja California, Mexico
Short-finned pilot whale (Globicephala macrorhynchus) aerial view, Baja California, Mexico
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Lemon shark pups (Negaprion brevirostris) spend the first 5-8 years of their life in mangrove forests. The tangle of roots provides protection from predators
Whale Shark (Rhincodon typus) and Golden trevally (Gnathanodon speciosus) Cenderawasih Bay, West Papua, Indonesia
Killer whale (Orcinus orca) hunting for herring fish (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Reef manta (Manta alfredi) mouth open feeding at night, South Ari atoll, Maldives, Indian Ocean
Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Aerial view of small coral island, Western Provinces, Solomon Islands
Common eider (Somateria mollissima), female on her nest with sea behind, Finland, May
Southern Rockhopper penguin (Eudyptes chrysocome) colony, Kidney Island, Falkland Islands, October. Vulnerable species
Silver Drummer (Kyphosus sydneyanus) fish off the coast of Neds Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Giant Manta Ray (Manta birostris), with Remoras (Echeneididae) Socorro Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March
Sea cucumber / Sea apple (Pseudocolochirus violaceus) Rinca, Komodo National Park, Indonesia
Tropical Flounder (Bothus mancus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Galapagos Shark (Charcharhinus galapagensis), Roca Partida islet, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Striped Dolphin (Stenella coeruleoalba) in the strait of Gibraltar, Spain
Nurse sharks (Ginglymostoma cirratum) over seagrass at night. White spiracle behind eye is visible. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Sea apple (Pseudocolochirus violaceus) Komodo National Park, Indonesia
Orange anemonefish (Amphiprion sandaracinos) in its host Mertens carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea
Flame anthias (Pseudanthias ignitus) displaying to the more drab female (above) during courtship in the evening. Similan Islands, Thailand. Andaman Sea, Indian Ocean
Great hammerhead shark (Sphyrna mokarran) female at night with Bar jacks (Caranx ruber), South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
School of Munks devil ray (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Fluorescent coral photographed with ultraviolet / UV light, Bonaire, Leeward Antilles, Caribbean region, Netherlands Antilles
Patagonian redfish (Sebastes oculatus) about 4cm, Comau Fjord, Patagonia, Chile
Mandarinfish (Synchiropus splendidus) Yap, Micronesia
Blue shark (Prionace glauca) seen from above with pilot fish, Azores Islands, Portugal, Atlantic Ocean
Dwarf lionfish (Dendrochirus brachypterus) opens its awingsa (in fact enlarged pectoralDwarf lionfish (Dendrochirus brachypterus) opens its wings (in fact enlarged pectoral fins) Seraya, Tulamben area, Bali, Indonesia, Java Sea
Bignose unicornfish (Naso vlamingii) profile of male displaying (the blue stripe at the bottom of the frame is the tail of another individual) Baa Atoll, Maldives. Indian Ocean
Chilean devil rays / Mobulas (Mobula tarapacana) offshore Santa Maria, Azores, Portugal, Atlantic Ocean
Plankton at night, Palau, Philippine Sea
