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RF - Reef manta (Mobula alfredi) female swimming close to a coral reef, while Cleaner wrasse (Labroides dimidiatus), tiny by comparison, pick parasites from its belly
RF - Portrait of Coral hermit crab (Paguritta harmsi) living in a hard coral colony. North Male Atoll, Maldives. Indian Ocean (This image may be licensed either as rights managed or royalty free.)
RF - A school of Crescent-tail bigeye (Priacanthus hamrur) on a coral reef. North Male Atoll, Maldives. Indian Ocean (This image may be licensed either as rights managed or royalty free.)
RF - Basking shark (Ceterhinus maximus) feeding on plankton in Senen Cove, Cornwall, UK. June. (This image may be licensed either as rights managed or royalty free.)
RF - Denise's pygmy seahorse (Hippocampus denise) looks out from its home in a sea fan (Annella sp.). Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Portrait of a Bigfin reef squid (Sepioteuthis lessoniana) in open water at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Portrait of a tarpon (Megalops atlanticus) on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Coral (Lobophytum sp. ) on a reef flat, with Hawksbill turtle (Eretmochelys imbricata)Coral (Lobophytum sp.) on a reef flat, with Hawksbill turtle (Eretmochelys imbricata) in background. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Blacktip reef shark (Carcharhinus melanopterus) swims through schools of fish. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Killer whales (Orcinus orca) at the surface in an Arctic fjord in Spildra, northern Norway, with sea birds. Arctic Ocean
Tawny nurse sharks (Nebris ferrungineus) at night. Vavuu Atoll, Maldives. Indian Ocean
Several mimic goatfish (Mulliodichthys mimicus) hide within a school of Bluestripe snapper (Lutjanus kasmira) on a coral reef. North Ari Atoll, Maldives. Indian Ocean
School of White collar butterflyfish (Chaetodon collare) pack together above a coral reef. North Male Atoll, Maldives. Indian Ocean
Orange painted frogfish (Antennarius pictus) fishing. Here the frogfish is illuminated by a snooted strobe and the packground sand lit with a red LED torch. Bitung, North Sulawesi, Indonesia
Thorny seahorse (Hippocampus histrix) gripping onto a sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Veined octopus (Amphioctopus marginatus) emerging from its home in an old clam shell. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Sleeping sperm whale, (Physeter macrocephalus) Dominica, Caribbean Sea, Atlantic Ocean
Nazca booby (Sula granti), pair on rocky shore at dawn. Punta Suarez, Espanola Island, Galapagos. May 2015
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Marine iguana (Amblyrhynchus cristatus), Punta Vicente Roca, Isabela Island, Galapagos
Flightless cormorant (Phalacrocorax harrisi) underwater, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador, April
White-vented storm petrel (Oceanites gracilis galapagoensis) walking on surface of water whilst foraging, Galapagos
Long-finned pilot whale (Globicephala melas) off Nova Scotia, Canada. October
Atlantic bluefin tuna (Thunnus thynnus) off Antigonish, Nova Scotia, Canada. October
Overhead view of a blue shark (Prionace glauca) off Halifax, Nova Scotia, Canada. July
A group of frilled anemones (Metridium senile) off Bonaventure Island in the Gulf of Saint Lawrence, Quebec, Canada. September
Head of a North Atlantic right whale (Eubalaena glacialis) showing callosities, pathches of roughened skin that are unique to each whale. Gulf of Saint Lawrence, Canada. IUCN Status: Endangered
North Atlantic right whale (Eubalaena glacialis) tail fluke, diving to feed in the Gulf of Saint Lawrence, Canada. Endangered species. July
Juvenile Pollock (Pollachius virens) school within the protective fronds of kelp near Port Joli, Nova Scotia, Canada. August
Aerial view of Carters Beach on the Southshore of Nova Scotia, Canada. August
Bottlenose dolphin (Tursiops truncatus) underwater playing in the shallows off Eleuthera, Bahamas
Green sea turtle (Chelonia mydas) resting in a coral reef. Eleuthera, Bahamas
Graysby fish (Cephalopholis cruentata) a type of grouper, hiding in a sponge off Eleuthera, Bahamas
Queen conch (Lobatus gigas) juvenile feeding on algae growing on Seagrass (Thalassia testudinum) blades. Eleuthera, Bahamas
A juvenile soldierfish, possibly a Crescent-tail bigeye (Priacanthus hamrur) in the open ocean at night off Anilao, Philippines
Larval flounder (possibly Laeops Kitahara) in the open ocean at night off Anilao, Philippines
Pygmy seahorse (Hippocampus bargibanti) holding on to a branch of a sea fan coral. Malapascua, Philippines
Sea Otter, (Enhydra lutris), young male, Elkhorn Slough, California, USA. January
Colonial anemones (Amphianthus nitidus) with Green urn sea squirts (Didemnum molle) (Atriolum robustum). Lembeh Strait, North Sulawesi, Indonesia
Thornback cowfish (Lactoria fornasini). Lembeh Strait, North Sulawesi, Indonesia
Boxer Crab (Lybia tessellata) with sea anemones in its claws for defense. Lembeh Strait, North Sulawesi, Indonesia
Potrait of a Cuckoo wrasse (Labrus bimaculatus) Loch Linnhe, west coast of Scotland, UK, September
Hermit crab (Pagurus bernhardus) on yellow sponge, Loch Etive, west coast of Scotland, UK, September
Seaweeds including Thongweed / Sea thong (Himanthalia elongata), Tangleweed kelp (Laminaria digitata), Toothed wrack (Fucus serratus) and Dulse (Palmaria palmata)
Rock archway at sunset, Isle of Lewis, Outer Hebrides, Scotland, UK, September 2014
Horse mussel (Modiolus modiolus) bed with Brittlestars (Ophiothrix fragilis) and Edible crab (Cancer pagurus), Shetland, Scotland, UK, September
Orange jewel anemones (Corynactis viridis) Sula Sgeir, Scotland, UK, February