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Atlantic sixgill shark (Hexanchus vitulus) swimming with open mouth. Cape Eleuthera, Bahamas
Close up at the dental plates of Three color parrotfish (Scarus tricolor) as it comes in for a bite of hard coral, Fiji
Banded sea krait (Laticauda colubrina) hunting in seagrass bed. Manado, North Sulawesi, Indonesia
Bonnethead shark (Sphyrna tiburo) hunting amongst fish in Turtlegrass (Thalassia testudinum) seagrass bed, view from below
Atlantic blue crab (Callinectes sapidus) hunting in Turtlegrass (Thalassia testudinum) seagrass bed, view towards sky. Florida Keys, Florida, USA
Orange and white soft corals, Dead mans fingers (Alcyonium digitatum) thriving beneath kelp (Laminaria hyperborea). St Abbs, Eyemouth, Berwickshire, Scotland, United Kingdom. North Sea
Cuttlefish (Sepia officinalis) male hovers over sea oak (Halidrys siliquosa). Babbacombe, Torquay, Devon, England, United Kingdom. English Channel. North East Atlantic Ocean
RF - Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and Sea plumes (Pseudopterogorgia sp)
Green sea turtle (Chelonia mydas) near surface in shallow water. The Bahamas
Caribbean reef squid (Sepioteuthis sepioidea) eating fish at night, portrait. The Bahamas
Spiny seahorse (Hippocampus guttulatus) adult female in a meadow of (Zostera marina) seagrass. Studland Bay, Dorset, England, United Kingdom. English Channel, North East Atlantic Ocean
White and yellow nudibranch (Polycera quadrilineata) feeding on a sea mat (Membranipora membranacea). Shetland Islands, Scotland, British Isles. United Kingdom. North East Atlantic Ocean
School of Oriental sweetlips (Plectorhinchus vittatus) gather tightly together as they rest during the day, with a Clown triggerfish (Balistoides conspicillum)
Black rockfish (Sebastes melanops) school shelter in a bull kelp forest (Nereocystis luetkeana). Race Rocks, Victoria, Vancouver Island, British Columbia, Canada
A portrait of a clown triggerfish (Balistoides conspicillum) on a coral reef. Misool, Raja Ampat, West Papua, Indonesia. Misool Marine Protected Area. Ceram Sea. Tropical West Pacific Ocean
A male pajama cardinalfish (Sphaeramia nematoptera) incubating eggs in his mouth (not clearly visible). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise
Tail of Blue whale (Balaenoptera musculus brevicauda). This may be the pygmy sub-species of blue whale, Balaenoptera musculus. Mirissa, Sri Lanka, Indian Ocean
Double exposure of a Tompot blenny (Parablennius gattorugine) with underwater sun beams. Swanage, Dorset, England, United Kingdom
A reef scene with an orange elephant ear sponge (Ianthella basta) and red sea whip (Ellisella sp) with schooling silversides fish above. Pelee Island, Misool, Raja Ampat, West Papua, Indonesia
Split level photo of mangrove scenery, with hard corals ( including Goniopora sp.; Heliopora sp; Porites sp.) growing below Red mangrove tree: (Rhizophora mangle)
Giant barrel sponge (Xestospongia testudinaria) releases a large cloud of gametes as it spawns on a coral reef, while a Threadfin hawkfish (Cirrhitichthys aprinus) looks
Portrait of an endemic Cuban hutia (Capromys pilorides) on a tropical beach. Jardines de la Reina, Gardens of the Queen National Park, Cuba
Black grouper (Mycteroperca bonaci) yawning on a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Cuban hutia (Capromys pilorides) stands up on its back legs on a tropical beach. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Endemic
Spinetail devil ray (Mobula mobular), four with rays of light and sparkle of sun on water, aerial view. Sea of Cortez, Baja California, Mexico. March
Humpback whale (Megaptera novaeangliae) spouting, rainbow effect created in spray, aerial view. Baja California, Mexico. March
RF - Open brain coral (Trachyphyllia geoffroyi) lives in symbiosis with zooxanthellae which are responsible for nutrition and the vibrant colours. Lembeh Strait, North Sulaweisi, Indonesia
Polar bear (Ursus maritimus) portrait, Franz Jozef Land, Arctic Russia. July
Monastery terraces on Skellig Michael, Skellig Islands World Heritage Site, County Kerry, Ireland, Europe. September 2015
Pacific white-sided dolphins (Lagenorhynchus obliquidens) underwater, Queen Charlotte Strait, British Columbia, Canada. September
Giant Pacific octopus (Enteroctopus dofleini) showing arms and suckers, Vernon Rock, Queen Charlotte Strait, British Columbia, Canada. September
A Giant rock scallop (Crassadoma gigantea) encrusted with and surrounded by several other invertebrates animals, including White anemones (Metridium sp.), White mushroom ascidians (Distaplia sp.)
Gooseneck barnacles (Pollicipes Polymerus) amongst kelp, Nakwakto Rapids, Slingsby Channel, British Columbia, Canada. September
Purple and Ochre sea stars (Pisaster ochraceus) preying on Pacific blue mussels (Mytilus trossulus). Also seen are Green Surf Anemones (Anthopleura xanthogrammica)
Sockeye salmon (Oncorhynchus nerka) males fighting in river Adams River, British Columbia, Canada. October
Bull kelp canopy (Nereocystis luetkeana) floating on the surface providing protective habitat for juvenile Rockfishes (Sebastes spp.) Browning Pass, Queen Charlotte Strait, British Columbia, Canada
Wolf eel (Anarrhichthys ocellatus) Croker Rock, Queen Charlotte Strait, British Columbia, Canada. September
Stubby squid (Rossia pacifica) Nigei Island, Queen Charlotte Strait, British Columbia, Canada
Opalescent nudibranchs (Hermissenda crassicornis) and Mushroom compound ascidians (Distaplia occidentalis). Browning Pass, Queen Charlotte Strait, British Columbia, Canada. September
Hooded nudibranchs (Melibe leonina) with oral hood for catching planktonic prey. In the Pacific Northwest they are often found clinging to bull kelp (Nereocystis luetkeana)
Sea butterfly (Cymbulia peronii) originally classed as Pseudothecosomata. Cymbuliidae are pelagic sea snails, marine gastropod molluscs. Atlantic Ocean, close to Cape Verde. Controlled conditions
Short-snouted seahorse (Hippocampus hippocampus)Tenerife, Canary Islands
Red-lipped blenny (Ophioblennius atlanticus) portrait, Tenerife, Canary Islands
Wide-eyed flounder (Bothus podas) portrait showing eyes above head, Tenerife, Canary Islands
Duckbill ray (Pteromylaeus bovinus) Tenerife, Canary Islands
Tiger moray (Enchelycore anatina) portrait, Tenerife, Canary Islands
Red starfish (Echinaster sepositus) group on sea floor, Tenerife, Canary Islands