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Humboldt squid (Dosidicus gigas) cannibalising another squid of the same species at night off Loreto, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Illuminated netdevil (Linophryne arborifera), female specimen, from deepsea Atlantic Ocean near Cape Verde at a depth of 1900m
Marbled electric ray (Torpedo marmorata) Bouley Bay, Jersey, British Channel Islands
Aggregation of thousands of Spider crabs (Leptomithrax gaimardii) for moulting, South Australia Basin, Australia. Pacific Ocean
Dwarf Minke whale (Balaenoptera acutorostrata) Great Barrier Reef, Australia. Coral Sea
Humpback whale (Megaptera novaeangliae) breaching, Prince William Sound, Alaska, July
Group of California sea lions (Zalophus californianus) in a kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Coloured rocks, Rackwick Bay, Isle of Hoy, Orkney Isles, Scotland, UK, October 2014
Harbour porpoise (Phocoena phocoena), Shetland, Scotland, UK, October
Grey seal (Halichoerus grypus) swimming over kelp forest, Shetland, Scotland, UK, August
Waves lashing the Four Lighthouse during winter storm, Northern Brittany, France, December 2011. Sequence 8/12
Orcas / Killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Group of California sea lions (Zalophus californianus) swimming in kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
European river otter (Lutra lutra) sleeping on rocky shore, Shetland, Scotland, UK, July
Looking down onf Clouds over Atlantic Ocean with rainbow colours
Humpback Whale (Megaptera novaeangliae) tail slapping, tail lob, lobtailing. Vancouver Island, British Columbia, Canada, July
Bottlenose dolphin (Tursiops truncatus) at surface, Moray Firth, Highlands, Scotland. May
European / Common Lobster (Homarus gammarus) in rocky retreat. Controlled conditions, Cornwall, UK. May
Grey seal (Halichoerus grypus) split level of young seal investigating the camera with its nose and whiskers, close to shore. Lundy Island, Devon, UK, Bristol Channel. August
White spotted eagle ray (Aetobatus narinari) feeding by digging in the sand and seagrass for invertebrates. East End, Grand Cayman, Cayman Islands, Caribbean Sea
Portrait of Blacktip grouper (Epinephelus fasciatus) resting in Barrel sponge (Xestospongia testudinaria) Mabul Island, Sabah, Borneo, Malaysia. Sulawesi Sea
Queen conch (Strombus gigas) extending its proboscis and stalked eyes at it moves across the sand by coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Lions mane jellyfish (Cyanea capillata) with feeding tentacles spread swimming in the sun in shallow water. Gulen, Norway. North East Atlantic Ocean
Great white shark (Carcharodon carcharias) from above. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
RF - Close up detail of the eye and skin pattern of a Giant hawkfish (Cirrhitus rivulatus). Socorro Island, Pacific Ocean
Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean
RF - Lemon shark (Negaprion brevirostris) swimming through sun rays in the late afternoon. Little Bahama Bank, Bahamas. West Atlantic Ocean
Male Great white shark (Carcharodon carcharias) with sunrays, Guadalupe Island, Mexico, Pacific Ocean
RF - Banggai cardinalfish (Pterapogon kauderni). Lembeh Strait, North Sulawesi, Indonesia
Oman anemonefish / clownfish (Amphiprion omanensis) in a a Bulb-tentacle sea anemone (Entacmaea quadricolor), coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Great hammerhead shark (Sphyrna mokarran) in shallow water. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
Blue and gold fusilier / Scissor tailed fusilier (Caesio caerulaurea) school swimming through shafts of sunlight, West Papua, Indonesia
Blacktip reef shark (Carcharhinus melanopterus) juvenile in shallow water, Maldives, Indian Ocean
Patters of sunlight on a shallow sandy seabed. North Sound, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Sargassum fish (Histrio histrio) near the surface on floating Sargassum seaweed, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Portuguese Man-of-War (Physalia physalis) close up of tentacles, Sargasso Sea, Bermuda
Detail of rhinopores and the colour pattern of a toxic Nudibranch (Goniobranchus coi) on a coral reef. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Silky shark (Carcharhinus falciformis), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba, January. Vulnerable
RF - California sea lion (Zalophus californianus) with mouth open close to the camera. The milk teeth of this pup show decay, which is quite usual. Los Islotes, La Paz, Baja California Sur, Mexico
California sea lion (Zalophus californianus) uses a Panamic cushion star (Pentaceraster cumingi) as a toy. The sealions pick up the starfish and then drop them and chase after them as they sink
Southern stingray (Dasyatis americana) female swimming over a shallow sand bank with ripples. Stingray City Sandbar, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs
Spotted eagle ray (Aetobatus narinari) school, Kagi kandhu, ; North Male Atoll, Maldives, Indian Ocean
Blue spotted trevally (Caranx bucculentus) hunting small fish in the reef, South Ari Atoll, Maldives, Indian Ocean
American crocodile (Crocodylus acutus), underwater, Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba, January. Vulnerable species
Great Blue shark (Prionace glauca) profile portrait viewed from slightly above, Pico Island, Azores, Portugal, Atlantic Ocean