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Ruffs displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Ruff displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Capercaillie (Tetrao urogallus) male displaying and calling at lek, Kamchatka, Far east Russia, May
Northern Cardinal (Cardinals cardinalis), male, perched during light snowfall, St. Charles, Illinois, USA, February, non-ex
Two wild Namib stallions fighting, Namib Nakluft National Park, Namib Desert, Namibia, September 2008
Blackbuck (Antilope cervicapra) two males, stand off at sunrise, Rajasthan, India
Blackbuck (Antilope cervicapra) two males fighting, Rajasthan, India
Male lion (Panthera leo) head portrait, Masai Mara, Kenya
Indian Leopard (Panthera pardus fusca), subspecies from Asia, male resting on tree, Nagarhole National Park, India
Tiger (Panthera tigris) male resting on road. Bandhavgarh National Park, India
Tiger (Panthera tigris) male in profile. Bandhavgarh National Park, India
Male Japanese giant salamander (Andrias japonicus) eating the eggs of another salamander, Japan, September
Stag Beetle (Lucanus cervus) in defensive posture; male in garden where it emerged naturally. Kent, UK, June. Controlled situation
Adult bull African Elephant (Loxononta africana) reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October
Bull Walrus (Odobenus rosmarus) in coastal waters off Spitsbergen. Arctic Norway, June
Regent bowerbird (Sericulus chrysocephalus) male perched, Lamington National Park, Queensland, Australia, October
Southern / Double-wattled cassowary (Casuarius casuarius) wild, adult male portrait, Atherton Tablelands, Queensland, Australia, November
Victorias riflebird (Ptiloris victoriae) male displaying in rainforest, Atherton Tablelands, Queensland, Australia, October
Buff breasted kingfisher / White tailed kingfisher (Tanysiptera sylvia) male, North Queensland, Australia
Wood Duck (Aix sponsa), male on water with autumn colour reflected in water, Ohio, USA
Prairie Warbler (Dendroica discolor) male singing on branch of tree, with spring leaves, Tompkins County, New York, USA, May
Male Eastern bluebird (Sialia sialis) feeding on winterberry (Ilex sp) fruit, New York, USA, January
Male American robin (Turdus migratorius) feeding on Winterberry (Ilex sp) New York, USA, January
Male Giant / Mammoth wasp (Megascolia flavifrons) close-up of face showing long antennae and small jaws, Stenje region, Galicica National Park, Macedonia, June 2009
Male Gyrfalcon (Falco rusticolus) in flight, Myvatn, Thingeyjarsyslur, Iceland, June 2009
Mediterranean Male Monk seal (Monachus monachus) Deserta Grande, Desertas Islands, Madeira, Portugal, August 2009, Critically endangered species
Large male Mediterranean Monk seal (Monachus monachus) Deserta Grande, Desertas Islands, Madeira, Portugal, August 2009, Critically endangered species
Male Yellow triplefin blenny (Tripterygion delaisi) Larvotto Marine Reserve, Monaco, Mediterranean Sea, July 2009
Male Longhorn beetle (Stenocorus sp) Queyras Natural Park, France, May 2009. WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS
Young male Red deer (Cervus elaphus) walking on snow field to get rid of flies, Western Tatras, Carpathian Mountains, Slovakia, June 2009
Red deer (Cervus elaphus) stag calling at sunset during rut, Klampenborg Dyrehaven, Denmark, September 2008
Male Stag beetle (Lucanus cervus) Suffolk, England, June 2009. WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS
Male Spanish luna / Isabelline moth (Graellsia isabellina) close-up of wings, France, Queyras Natural Park, May 2009. WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS
Male Chaffinch (Fringilla coelebs) on branch, Los Tilos National Park, La Palma, Canary Islands, Spain, March 2009
Male Western green lizard (Lacerta bilineata) warming up, San Marino, May 2009. WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS
Roe deer (Capreolus capreolus) male amongst reeds in marsh, Matsalu National Park, Estonia, May 2009
Marsh warbler (Acrocephalus palustris) male singing at dawn, Lithuania, June 2009
Wild Iberian lynx (Lynx pardinus) male carrying rabbit prey, Sierra de Andjar Natural Park, Mediterranean woodland of Sierra Morena, north east Jan Province, Andalusia, Spain, May 2009
Wild Iberian lynx (Lynx pardinus) male walking, Sierra de Andjar Natural Park, Mediterranean woodland of Sierra Morena, north east Jan Province, Andalusia, Spain, May 2009, Critically endangered
Marsh frog (Rana ridibunda) calling, Sierra de Andjar Natural Park, Mediterranean woodland of Sierra Morena, north east Jan Province, Andalusia, Spain, April 2009
Wild Iberian lynx (Lynx pardinus) male, Sierra de Andjar Natural Park, Mediterranean woodland of Sierra Morena, north east Jan Province, Andalusia, Spain, May 2009, Critically endangered
Wild Iberian lynx (Lynx pardinus) male, one year, with GPS tracking collar, sitting, Sierra de Andjar Natural Park, Mediterranean woodland of Sierra Morena, north east Jan Province, Andalusia, Spain
Roe deer (Capreolus capreolus) buck in wet meadow, Nemunas Regional Park, Lithuania, June 2009
Whinchat (Saxicola saxicola) male singing perched on post, Lithuania, May 2009
Male Whinchat (Saxicola saxicola) perched, Lithuania, May 2009
Two male Saiga antelopes (Saiga tatarica) Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Male Saiga antelope (Saiga tatarica) silhouetted, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Great reed warbler (Acrocephalus arundinaceus) perched on reed, singing, Eastern Slowakia, Europe, June 2009