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Ruff (Philomachus pugnax) male displaying, Varanger, Norway, May
Polar bear (Ursus maritimus) young male playing / rolling around in the snow along a barrier island during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, Alaska
RF- Rear view of male Bengal tiger (Panthera tigris tigris) walking on track. Bandhavgarh National Park, India. Endangered species
Dusky dolphins (Lagenorhynchus obscurus) courting pair leaping from the sea, Kaikoura, Canterbury, New Zealand, January
Arctic fox male portrait {Vulpes lagopus} Norway captive
Kirks Dik-Dik (Madoqua kirki) male standing in tall grass, Tarangire National Park, Tanzania
Male Indian peafowl / peacock (Pavo cristatus) from open forest areas on the Indian subcontinent. Captive, Jurong Bird Park, Singapore
RF- Adult male Serval (Felis serval) stalking prey in long grass. Serengeti National Park, Tanzania
Male Black and white colobus monkey {Colobus guereza} Lake Naivasha, Kenya, East Africa
Blue-chested hummingbird (Amazilia amabilis) male feeding from flower, Costa Rica, Central America
Coppery-headed Emerald, Elvira cupreiceps, male in flight feeding on Spiral Ginger (Costus Pulverulentus), Central Valley, Costa Rica, Central America, December 2006
Male White-throated Mountain Gem (Lampornis castaneoventris). Tamalanca Mountains, Costa Rica. May
Takin male portrait {Budorcas taxicolor} Mt Qinling, Shaanxi, China
Kingfisher (Alcedo atthis) male perched in tree with mud on beak, Hertfordshire, England, UK, March
Bullocks Oriole (Icterus gabula bullockii) male in breeding plumage perched amid new leaves in spring, Mono Lake Basin, California, USA
Eastern Bluebird (Sialia sialis) male perched on flowering eastern redbud in spring, New York, USA (Digitally retouched image - flower cleanup)
Bullfinch (Pyrrhula pyrrhula) male perched on frosty twigs, Kuusamo, Finland, January
Lesser Goldfinch {Carduelis psaltria} black-backed male on Mealy sage (Salvia farinacea) Hill Country, Texas, USA
Rear view of male Bengal tiger walking {Panthera tigris tigris} Kanha NP, India - Winner in Wildlife Photographer of Year competition
California Quail (Callipepla californica), adult male perched on bush, acting as sentinel, Montana De Oro State Park, California, USA, July
Common cranes displaying by woodland {Grus grus} Finland
Male Reindeer (Rangifer tarandus) on autumn tundra. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, October
Greenfinch (Carduelis chloris) on blackthorn / sloe berries, UK
Red deer (Cervus elaphus) stag roaring during the rut, Leicestershire, UK, October. Non-ex
Mallard drake (Anas platyrhynchos) portrait of male on water, with autumn colours reflecting. Lancashire, England, UK. November
RF- Northern Cardinal (Cardinalis cardinalis) immature male perched on Texas Red Oak (Quercus Texana). New Braunfels, San Antonio, Hill Country, Central Texas, USA
Moose (Alces alces) bull at the edge of the Great North Woods, Baxter State Park, Maine, New England, USA, Autumn
Male Greenfinch (Carduelis chloris) perching on Hawthorn, Cheshire, UK, October
Red deer {Cervus elaphus} adult male in autumnal bracken during the October rut. Leicestershire, UK
Red deer stag {Cervus elaphus} calling in the mist, Dyrehaven, Denmark
Roe deer (Capreolus capreolus) buck feeding in a field at dusk. Perthshire, Scotland, August
Red deer stag (Cervus elaphus) portrait lying in long grass, Richmond Park, London, UK, August
Migrant Hawker dragonfly (Aeshna mixta latrielle) male in flight, Wiltshire, England. August
Male Keeled skimmer (Orthetrum coerulescens) in flight, Holt Lows CP, Norfolk, England, August
Black-throated Green Warbler (Dendroica virens), male in breeding plumage singing from hemlock branch, New York, USA
Male Spanish ibex (Capra pyrenaica) Sierra de Gredos, Spain, November 2008
Cuckoo wrasse (Labrus mixtus) male profile, Channel Islands, UK August
Male African lion (Panthera leo), Kruger National Park, Transvaal, South Africa, September
Orangutan (Pongo pygmaeus) adult male, Nyaru Menteng Orangutan Reintroduction Project, Central Kalimantan, Borneo, Indonesia
Orang utan (Pongo pygmaeus) head portrait of dominant male Richie, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Male Jaguar through vegetation {Panthera onca} captive Pantanal, Brazil
Male Giant panda (Ailuropoda melanoleuca) portrait, lying on snow, approximately 10 years, captive
Ruffs displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Ruff displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Capercaillie (Tetrao urogallus) male displaying and calling at lek, Kamchatka, Far east Russia, May
Northern Cardinal (Cardinals cardinalis), male, perched during light snowfall, St. Charles, Illinois, USA, February, non-ex
Two wild Namib stallions fighting, Namib Nakluft National Park, Namib Desert, Namibia, September 2008
Blackbuck (Antilope cervicapra) two males, stand off at sunrise, Rajasthan, India
