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Rio Jatuntianhua Robber Frog (Pristimantis eriphus) male on a leaf. Ecuador, Vulnerable species
Tabby kittens, Stanley and Fosset, 12 weeks old, walking together
Masai Giraffe (Giraffa camelopardalis tippelskirchi) male and female (back). Masai-Mara Game Reserve, Kenya
Bluethroat (Luscinia svecica) male singing, Sinirinta, Kuusamo, Finland, June. Fascinating birds bookplate
Alpine ibex (Capra ibex) adult male chasing two young males away, Gran Paradiso National Park, Italy, July
Andalusian stallion running out of the sea on beach, Ojai, California, USA
Wild horses / Mustangs, two stallions play fighting, Pryor Mountains, Montana, USA, July 2010
African Lion (Panthera leo) young male at sunrise, Etosha National Park, Namibia
A rare East Bulgarian stallion cantering at the Kabiuk National Stud, Shumen, Bulgaria, February 2012
Red deer (Cervus elaphus) stags fighting at dusk during rutting season, Cheshire, UK October
Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA
Domestic rabbit, French Lop, Blue, young male, 6 months, studio portrait
Mustang / wild horses, second year cremello colt Claro play fighting with bay, McCullough Peak herd, Wyoming, USA, July 2009
Rock beauty angelfish (Holacanthus tricolor) pair at dusk performing their courtship dance with the larger male circling the female, Georgetown, Grand Cayman, Cayman Islands, British West Indies
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
Grizzly Bear (Ursus arctos horribilis) male in water. Coastal Katmai National Park, southwest Alaska, USA, August
Male African Lion (Panthera leo) playing with 4 month cub. Big Marsh, near Ndutu, Nogorongoro Conservation Area, Serengeti National Park, Tanzania
Male Greenfinches (Carduelis chloris) squabbling in flight. Dorset, UK, March
Green crowned brilliant hummingbird (Heliodoxa jacula) male taking off to feed at feeder in garden, another waiting perched on lichen covered branch, Costa Rica, January
Emperor penguins (Aptenodytes forsteri) males dive through openings in the sea ice to take their first swim in over 4 months as they have spent the bleak winter incubating their eggs
Chestnut arabian stallion running, California, USA
Grey Andalusian / Spanish stallion running, California, USA
Male Andean Cock-of-the-Rock (Rupicola peruvianus) displaying. Cock-of-the-Rock Lodge, Peru, August
Male Panther Chameleon (Furcifer pardalis) feeding on Preying mantis in beach side vegetation. Bay of Antongil, Masoala Peninsula National Park, north east Madagascar
European Mouflon (Ovis musimon) male head profile. The Netherlands, January
British Shorthair tomcat, blue coated, portrait standing in straw with Pumpkins / Squash
Large adult male Leopard (Panthera pardus) walking through the bush at night, Sabi Sand Private Game Reserve, South Africa, June
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male calling and performing courtship / territorial display in cattail marsh, Mono Lake Basin, California, USA
Great Dane, black coated bitch, and Magyar Vizsla / Hungarian Pointer, tan coated maleGreat Dane, black coated bitch, and Magyar Vizsla /Hungarian Pointer, tan coated male, double portrait in profile together
Violet Sabrewing Hummingbird (Campylopterus hemileucurus) male hovering at feeder, Bosque de Paz, Costa Rica, Central America
African Elephant (Loxodonta africana) bulls in dominance fight, Mono Pools national Park, Zimbabwe, Southern Africa
A Shagya Arab stallion (Equus caballlus) is shown in hand in the courtyard of the Babolna Arabian Stud, Babolna, Komarom-Esztergom, Hungary
Portrait of male Boxer, light brindle coloured with natural ears, standing by lake shore, Illinois, USA
Weddell seal (Leptonychotes weddellii) male surfacing in ice hole, McMurdo Sound, Ross Sea, Antarctica, November 2008
Male Comet / Madagascar moon moth (Argema mittrei) recently emerged drying its wings. Andasibe-Mantadia National Park, Madagascar
Green woodpecker {Picus viridis} male in flight, Lorraine, France
Trumpeter Swans (Cygnus buccinator) in winter morning mist, showing aggression during courtship behaviour, Mississippi River, Minnesota, USA, February
Male Atlantic salmon {Salmo salar} leaping, migrating upstream to spawn, Perthshire, Scotland, UK, October
Red Bird of Paradise (Paradisaea rubra) adult male at lek display site in the top of a canopy tree. Batanta Island, Papua, Indonesia
Grey Andalusian stallion cantering with Rocky mtns behind, Colorado, USA
Gelada baboon mature male portrait with wind blown hair (Theropithecus gelada) Simien Mountains NP, Ethiopia
Top of male Lions head {Panthera leo} Selous Game Reserve, Tanzania
Male Lion walking towards camera {Panthera leo} Masai Mara, Kenya
Two male Ostriches (Struthio camelus) running during dispute, Etosha NP, Namibia
African Lion (Panthera leo) cub reaches for a moment of intimacy with its father, Northern Tuli Game Reserve, Botswana
Wild Chincoteague (Equus caballus) two breeding stallions greeting one another, Chincoteague National Wildlife Refuge, Chincoteague Island, Virginia, USA, June
Mallards (Anas platyrhynchos) slipping on ice on frozen lake, Wiltshire, UK, January
Ruff (Philomachus pugnax) male displaying, Varanger, Norway, May
