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Coalition of male Lions (Panthera leo) in grassland before storm, Masai-Mara game reserve, Kenya
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
Four Cedar waxwings (Bombycilla cedrorum) feeding on Crabapple (Malus sp) fruit, Ithaca, New York, USA, February
Masai Giraffe (Giraffa camelopardalis tippelskirchi) juveniles, Masai Mara Game Reserve, Kenya
Vancouver Island black bear (Ursus americanus vancouveri) mother with cubs on a beach, Vancouver Island, British Columbia, Canada, July
American Red fox (Vulpes vulpes fulva) baby leaping on its disinterested mother. Grand Teton National Park, Wyoming, USA, May
Northern raccoon (Procyon lotor), group standing on branch, captive
Lionesses (Panthera leo) grooming each other, Masai-Mara Game Reserve, Kenya
Rural view of rolling countryside with grazing cattle, near Frome, Somerset, UK. July 2014
Roseate Spoonbills (Platalea ajaja) in flight, Pantanal, Mato Grosso do Sul, Brazil
Hyacinth Macaw (Anodorhynchus hyacinthinus) small group in flight, Brazil
Mute Swans (Cygnus olor) foraging, southwest Finland, February
Savanna landscape wirh Grants zebras (Equus quagga boehmi) and acacia trees, Masai Mara National Reserve, Kenya, Africa
Liver Flatcoated Retriever puppy, 6 weeks, with two baby Guinea pigs
Fallow deer (Dama dama) stags, captive, Cabarceno Park, Cantabria, Spain, June
Black bears (Ursus americanus) three cubs, one climbing, Minnesota, USA, June
Atlantic Puffins (Fratercula arctica) three resting on a snow bank, Hornoya, Varanger, Norway, April
Appennine Trumpet Gentian (Gentiana dinarica) in flower, on roadside below Gran Sasso, Appennines, Abruzzo, Italy, May
Three Llamas (Lama glama) resting in the Machu Picchu ruins. Urubamba, Peru. August, 2016
Southern stingrays (Hypanus americanus) swimming over a sand bar in the early morning. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Four California spiny lobsters (Panulirus interruptus) shelter beneath a boulder in a kelp forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Sunflowers (Helianthus annuus) in bloom, Norfolk, England, UK, August
Imbabura Tree Frogs (Boana picturatus) perched on a branch. Bilsa, Esmeraldas, Ecuador
Monte Perdido massif and Las Tres Sorores (The Three Sisters) Ordesa National Park, Aragon, Huesca, Spain
Mute swan (Cygnus olor) cygnets on water, Lac de Saint Point, Franche Comte, France, May
Plains zebra (Equus burchelli) three on dried mud plains, Etosha National Park, Namibia
Magnificent bird of paradise (Cicinnurus magnificus) male, leaning back and presenting his breast shield and raised cape to two females, courtship display, Arfak Mountains, West Papua, Indonesia
Adelie penguins (Pygoscelis adeliae) on large iceberg with towers and pinnacles in background, Weddell Sea, Antarctica
Four Common / Plains Zebras (Equus quagga) standing side by side, portrait, Okavango Delta, Botswana, Africa
African wild dog (Lycaon pictus) alpha female, interacting with two of her pups, aged 4 weeks, Okavango Delta, Northern Botswana, Africa
Three Brown noddies / Common noddies (Anous stolidus) trying to steal a fish from a Brown pelican (Pelecanus occidentalis), Santiago Island, Galapagos Islands, Pacific Ocean
Three King penguins (Aptenodytes patagonicus) reflections in water, Salisbury Plain, South Georgia, Southern Ocean
Common blue butterflies (Polyommatus icarus) roosting in morning light, Devon, UK. July
Puma (Puma concolor) female with two cubs, aged six months, sitting in fresh snow, Torres del Paine National Park, Patagonia, Chile
Three Spotted hyenas (Crocuta crocuta) fighting over meat from a nearby carcass with flock of White-backed vultures (Gyps africanus) watching in background, Mana Pools National Park, Zimbabwe
Adult Guanacos (Lama guanicoe) grazing in deep snow near Lago Pehoe, Torres del Paine National Park, Chile, July
Giraffe (Giraffa camelopardalis angolensis) calf, aged 6 weeks, with two Yellow-billed oxpeckers (Buphagus africanus) perched on its head feeding on parasites, Okavango Delta, Botswana, Africa
American flamingos (Phoenicopterus ruber) wading through salt pond, Quinta Playa, Isabela Island, Galapagos Islands, Ecuador
Three male Ostriches (Struthio camelus) running and flapping wings in aggressive display, Kgalagadi Transfrontier Park, South Africa
Female Cheetah (Acinonyx jubatus) with juveniles crossing grassland in rain storm, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Four White storks (Ciconia ciconia) perched on street lights, silhouetted at dusk, Madrid, Spain. June
Fimbriated moray / Darkspotted moray (Gymnothorax fimbriatus) peering out from crevice with two Banded coral shrimp (Stenopus hispidus) close by, Philippines, Pacific Ocean
Four Grey partridges (Perdix perdix) on snow-covered arable field, Scottish Borders, Scotland, UK. January
Female polar bear (Ursus maritimus) with two cubs walking in a line across snow, Churchill, Canada. November
Bison (Bison bison) being circled by wolves (Canis lupus) hunting in snow, Yellowstone National Park, USA. January
Four Wolves (Canis lupus) walking in snow, Yellowstone National Park, USA. January
RF - Group of four Mycena toadstools on mossy stump. The New Forest, Hampshire, UK. October. (This image may be licensed either as rights managed or royalty free)
Three female Putorana snow sheep (Ovis nivicola borealis) climbing up rocky mountainside with lamb following, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia