Goldfinch (Carduelis carduelis) perched on twig in garden hedge in winter, Scottish Borders, Scotland, December
Bison (Bison bison) being circled by wolves (Canis lupus) hunting in snow, Yellowstone National Park, USA. January
Striped marlin (Tetrapturus audax) feeding on Sardine bait ball (Sardinops sagax). West Coast of Baja California Peninsula, Mexico. Pacific Ocean
Santa Catalina Island rattlesnake (Crotalus catalinensis), only rattlesnake without rattle, slithering in rock. Santa Catalina Island, Loreto Bay National Park, Sea of Cortez, Mexico. May
Female Serval (Leptailurus serval) portrait. Masai Mara national reserve, Kenya, East Africa
Curlews (Numenius arquata) group flying over the sea during storm. Wales, UK December. Highly commended, Habitat division, British Wildlife Photography Awards (BWPA) competition 2012
Two donkeys grazing on grassland to help with conservation, Carry Farm, Argyll, Scotland, UK. August