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Polar bear (Ursus maritimus) female and cubs, one cub walking up slope of ice in foreground. Svalbard, Norway, July
Mute swan (Cygnus olor) swimming with cygnets in misty lake in morning light. Richmond Park, London, England, UK. May. Cropped image
American quarter horse, three standing in snow. Alberta, Canada. February
Eastern white-bearded wildebeest (Connochaetes taurinus), four running in line, on migration, Masai Mara National Reserve, Kenya
Snowdrop (Galanthus nivalis) anthers and inner petals. Outer petals have been removedSnowdrop (Galanthus nivalis) anthers and inner petals.Outer petals have been removed. Focus stacked
Common rue (Ruta graveolens) flower in visible light, nectar at base of petals. Stamens flip up sequentially with one still folded over the receptacle. Surrey, England, UK
Common rue (Ruta graveolens) flower in UV light, pollen and nectar at base of petals fluorescing. Insect pollinators are able to see position of rewards on offer. Surrey, England, UK
Porcelain orchid (Chloraea magellanica) with osmophores on lip fluorescing in UV light. Chile. Controlled conditions, focus stacked. Sequence 2/2
Trailing fuschsia (Fuchsia procumbens) cultivated in alpine house, Surrey, England, UK. Native to New Zealand
Great horned owl (Bubo virginianus) preening two chicks in nest in Saguaro (Carnegiea gigantea) cactus, Sonoran desert, Arizona, USA
Scarlet Macaws (Ara macao) drinking water inside an Amazonian claylick. Yasuni National Park, Orellana, Ecuador
People watching the Fissure eruption of Chico volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005
Lesser horseshoe bats (Rhinolophus hipposideros) in magnesium mine, Shropshire, England, UK, April
English oak (Quercus robur) ripening acorns, Dorset, UK September
Wirehaired daschund, Staffordshire bull terrier and Hungarian viszla in autumn colours, Wiltshire, UK
Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions
Red-breasted Mergansers (Mergus serrator) male in foreground performing courtship display to female in background (left), Aurora, New York, USA, April
Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October
Black-bellied whistling-ducks (Dendrocygna autumnalis), group perched amid Spanish moss on bald cypress tree, Lakeland, Florida, USA, January
Guanaco (Lama guanicoe) grazing on grassland plain, Patagonia, Chile. Novmeber
Three French Bulldog puppies, age 6 weeks, sitting in a row
Green-crowned brilliant (Heliodoxa jacula), three feeding on Lobster-claw (Heliconia sp) flower in rain. Costa Rica
Thomsons gazelle (Eudorcas thomsonii) male and two females standing in rainstorm, Masai Mara Game Reserve, Kenya
Blackcap (Sylvia atricapilla) female and two males huddling together for warmth before going to sleep, Finland, May
Gentoo penguin (Pygoscelis papua) walking back to the colony, Sea Lion Island, Falkland Islands, October
Eastern Ring-tailed Vontsira (Galidia elegans elegans) (endemic family Eupleridae) foraging on forest floor. Marojejy National Park, north east Madagascar
Starling (Sturnus vulgaris), eating fruit of Japanese Persimmon (Diospyros kaki), Provence, France
Series of Bald eagle (Haliaeetus leucocephalus) in flight, Homer, Alaska, USA, March. Digital composite
Polar bear (Ursus maritimus) sow with two cubs walking past a pile of Bowhead whale (Balaena mysticetus) bones, Kaktovik, Barter Island, 1002 Area, Arctic National Wildlife Refuge, North Slope
Chilean devil rays / Mobulas (Mobula tarapacana) offshore Santa Maria, Azores, Portugal, Atlantic Ocean
Alpine ibex (Capra ibex) group posing on a rocky mountain side on misty day, Gran Paradiso National Park, Italy. November
Banded demoiselle (Calopteryx splendens) males resting on grass, Netherlands, August
RF- Domestic pig, hybrid large white sow and piglets in sty, UK, September 2010
Antarctic fur seal (Arctocephalus gazella), mother warning off curious juvenile. Southern elephant seal (Mirounga leonina) in background. Gold Harbour, South Georgia. October
Razorbill (Alca torda) group, Great Saltee Island, County Wexford, Ireland. June
Candle-snuff fungus (Xylaria hypoxylon), New Forest National Park, Hampshire, England, UK. November
Bornean orangutan (Pongo pygmaeus) mother and babies, Tanjung Puting National Park, Borneo-Kalimatan, Indonesia, endangered species
RF - Leopard (Panthera pardus) mother with cubs, Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Arctic fox (Alopex lagopus) juveniles play fighting, Wrangel Island, Far East Russia, August
Musk ox (Ovibos moschatus) herd, Wrangel Island, Far Eastern Russia, March
Split level view of Brown pelicans (Pelecanus occidentalis) feeding, Tortuga Bay, Santa Cruz Island, Galapagos
Pallass cat (Otocolobus manul) three cats, Tibetan Plateau, Qinghai, China
Row of three trees in snow, Groot Schietveld, Wuustwezel, Belgium, January 2010
American buffalo (Bison bison) with group of calves, Yellowstone National Park, Wyoming, USA. May
Chilean Flamingo, Phoenicopterus chilensis, La Pampa, Argentina
A family of otters rest on the intertidal seaweed. European river otter (Lutra lutra) Shetland, Scotland, UK, July
Great crested grebe (Podiceps cristatus) close-up of an adult with two young chicks. The Netherlands. June 2014
White waterlily (Nymphaea alba) in a peat bog lake, De Weerribben-Wieden National Park, Holland. September