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Three female Putorana snow sheep (Ovis nivicola borealis) climbing up rocky mountainside with lamb following, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Three Exmoor ponies (Equus ferus caballus), semi-feral native breed, in Exmoor National Park, Somerset / Devon, England. November
Alpha male Grey wolf (Canis lupus) biting angry young wolf to assert dominance, Finland. July
Small flock of migrating Black-tailed godwits (Limosa limosa) in flight against a bright blue sky, River Coquet Estuary, Northumberland, UK. September
RF - Eschmeyers scorpionfish (Rhinopias eschmeyeri) composite image showing different colour variations on black background. Lembeh Strait, North Sulawesi, Indonesia
Three Lined seahorses (Hippocampus erectus), two male and one female, feeding on plankton in an alkaline saltwater pond, The Bahamas, Caribbean
Blue pincushion (Brunonia australis). Tasmania, Australia. December
Dominant male Chimpanzee (Pan troglodytes troglodytes) walking in the mangrove, with female carrying her infant, Conkouati-Douli National Park, Republic of Congo, Africa
Juvenile Mute swan (Cygnus olor) in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. August
Three Grey partridge (Perdix perdix) walking in snow. Near Nijmegen, the Netherlands. February
Dominant male Chimpanzee (Pan troglodytes troglodytes) in the mangrove, with female and infant vocalising, Conkouati-Douli National Park, Republic of Congo, Africa
Female Galapagos sea lion (Zalophus wollebaeki), sleeping together, likely to be related, Santa Fe Island, Galapagos, South America
Galapagos penguin (Spheniscus mendiculus) roosting on rocks in late afternoon, Isabela Island, Galapagos, South America
Silvery grebe (Podiceps occipitalis) attempting to join Hooded grebes (Podiceps gallardoi) during a breeding display, Patagonia National Park, Santa Cruz, Argentina. Cropped
Four Hereford bullocks looking over a fence with Cow Parsley (Anthriscus sylvestris) Worcestershire, England, May
Three Mallards (Anas platyrhynchos) on shallow pond on misty morning at sunrise, Valkenhorst Nature Reserve, The Netherlands, September
Three Spoonbills (Platalea leucorodia) feeding in shallow pond, Valkenhorst Nature Reserve, The Netherlands, August
Whooper swans (Cygnus cygnus) in flight with cityscape behind, Sanmenxia, Henan province, China
Common spotted orchids (Dactylorhiza fuchsii), Kingcombe Meadows, Dorset, UK, June
Pilot whale (Globicephala macrorhynchus) pod, Tenerife, Canary Islands. August
Fiordland penguins (Eudyptes pachyrhynchus) walk along a forest path, heading back to their burrows, after foraging in the ocean. North of Hst, Westland District, South Island, New Zealand
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
European Bison (Bison bonasus) in winter, Białowieza National Park, Poland. January
Red stingray (Hemitrygon akajei), four in a row, viewed from below. Chiba Prefecture, Honshu, Japan. October
Himalayan griffon vulture (Gyps himalayensis) Angsai Nature Reserve (Valley of the Cats), Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China
Green turtle (Chelonia mydas) resting on sea floor and two juvenile Golden trevally (Gnathanodon speciosus). Marsa El Shouna, Marsa Alam, Egypt
Highland cows, three silhouetted at dusk. Glengorm Estate, Isle of Mull, Scotland, UK. June
RF - North American river otters (Lontra canadensis) (probably female with two juveniles) on the frozen river edge.Yellowstone National Park, USA. January. Digitally stitched image
Group of Longfin batfish (Platax teira) beneath the surface, close to an island. Farondi Islands, Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Steller sea lions (Eumetopias jubatus) swimming in the Bering Sea near Verkhoturova Island, Russia
Mute swan (Cygnus olor) in a threathening territorial pose and attacking a rival Lake Geneva, near Geneva, Switzerland, March
Bewicks swan (Cygnus columbianus bewickii) adults swimming on a marshland pool, Gloucestershire, UK, November
Waved albatrosses (Phoebastria irrorata) courting, Punta Suarez, Espanola Island, Galapagos
Caribbean Reef Shark (Carcharhinus perezi) group of three, Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba
Three common blue damselflies (Enallagma cyathigerum) roosting on sedge and silhouetted at dawn, Broxwater, Cornwall, UK. July
RF- Greater flamingo (Phoenicopterus roseus) group of three silhouetted at sunset, Pont Du Gau Park, Camargue, France. (This image may be licensed either as rights managed or royalty free.)
Galah (Cacatua roseicapilla) in tree, Piccaninny Plains. Cape York Peninsula, Queensland, Australia
Little tern (Sterna albifrons) three eggs in nest scrape, County Wicklow, Ireland, June
Coyote (Canis latrans) foraging in deep winter snow disturbed by grazing American bison (Bison bison) Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Whooper Swans (Cygnus cygnus) in snow. Japan, February
Three Marbled white butterflies (Melanargia galathea) nectaring on Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June
Blue tit (Cyanistes caeruleus) Sierra de Grazalema Natural Park. Southern Spain. July
Four Pygmy seahorses (Hippocampus bargibanti) on a Seafan / Gorgonian (Muricella sp. ) including one or several pregnant males. New Caledonia, Pacific Ocean
Crab-plover (Dromas ardeola) three flying, two with crabs in beaks. Oman, June
RF - Cape buffalo (Syncerus caffer) drinking at night, Zimanga private game reserve, South Africa. (This image may be licensed either as rights managed or royalty free.)
Rocky mountain bighorn sheep (Ovis canadensis canadensis) grazing. Lamar Valley, Yellowstone National Park, Wyoming, USA. January
Southern right whales (Eubalaena australis) aerial view of three adullts engaged in social activity, photographed with the permission of the Department of Environmental Affairs, South Africa
RF - King penguin (Aptenodytes patagonicus), four returning to sea. St Andrews Bay, South Georgia. November. (This image may be licensed either as rights managed or royalty free.)
