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A female Spiny Seahorse (Hippocampus guttulatus) in weeds. Sardina, Gran Canaria, Canary Islands, East Atlantic Ocean, October
In-camera motion blur image of a female Yellow Seahorse (Hippocampus kuda). Lembeh Strait, Sulawesi, Indonesia, Molluca Sea, March
RF- Orange Tip butterfly (Anthocharis cardamines) female resting on Cuckooflower (Cardamine pratensis). North Devon, UK. April
Female Azure damselfly (Coenagrion puella), Vallee de la Seille (Seille Valley), Lorraine, France, June
RF - Black panther / melanistic Leopard (Panthera pardus) peering through leaves, captive
Asiatic lionesses and cubs (Panthera leo persica) drinking from pool, Gir Forest NP, Gujarat, India. Water is scarce in the Gir
Hanuman Langur / Northern Plains Grey Langur (Semnopithecus entellus) mother with baby in tree. Kanha National Park, Madhya Pradesh, India
Sperm Whale (Physeter macrocephalus) calf swimming uder its mother. Sao Miguel Island, Azores, July
Domestic Sheep (Ovis aries) ewe and two lambs, probably Romney x Perendale. Unshorn. Cape Kidnappers, Hawkes Bay, New Zealand, September
Portrait of a Lynx (Lynx lynx) mother and kitten. Hanau, Germany, July. Captive
Snow goose with gosling (Chen caerulescens) at sunset. Taken on location for BBC Frozen Planet series, 2008
Wild / feral Dulmen ponies (Equus caballus) close up of herd of mares and foals running on the Duke of Croys estate, Meerfelder Bruch, North Rhine-Westphalia, Germany, May 2011
Grey seal (Halichoerus grypus) adult female among breaking waves, Lincolnshire, UK Non-ex
Border Collie bitch, running towards the camera
Chinchilla Persian female cat, 6 years
Black-and-white Border Collie bitch lying chin on floor with black-and-white kitten
Border Collie with one of her puppies, 5 weeks, yawning
Black Swallowtail Butterfly (Papilio polyxenes), female on Cowpen Daisy / Golden Crownbeard (Verbesina encelioides). Laredo, Webb County, Texas, USA, April
Wild horse / mustang, mare rolling, Pryor Mountains, Montana, USA, July
Young Tigress (Panthera tigris), 22 month, being frightened by a dead leaf. Banghavgarh National Park, India, April
Female Rocky Mountain bighorn sheep ewe (Ovis canadensis canadensis) warning her newborn lamb that a rocky ledge is too close and dangerous
Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January
Proboscis Monkey (Nasalis larvatus) female sitting in tree, partially obscured by soft focus leaves in foreground, Bako National Park, Sarawak, Borneo, Malaysia
Polar bears (Ursus maritimus) female with cubs aged two years travelling along a barrier island during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, Alaska
An adult female California sea lion (Zalophus californianus) head portrait, in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
Crested Newt (Triturus cristatus carnifex) underwater view of female swimming on surface, Traversari lake, Campigna National Park, Emilia Romagna and Tuscany regions, Italy
Jaguar (Panthera onca) female, hiding in dense high grass. Cuiaba river, Pantanal, Mato Grosso do Sul Province. Brazil, South America
A wild Misaki-uma breeding stallion (right) brings back one of his mares to his band while another breeding stallion (left) charges him away, in the Cape Toi Reserve, Miyazaki Prefecture
African lion (Panthera leo) male and female at sunrise, Masai Mara National Reserve, Kenya, January
Humpback whale (Megaptera novaeangliae) mother and calf swimming, Hawaii, Pacific Ocean
Blue dacnis (Dacnis cayana) female visiting Fruiting shrub (Miconia longifolia) La Selva Field Station, Costa Rica
Ring tailed lemur (Lemur catta) mother and very young (1-2 week) baby. Berenty Private Reserve, Madagascar
Bengal tiger (Panthera tigris) tigress Arrowhead running through water, Ranthambhore, India