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Kelp greenling (Hexagrammos decagrammus), female, resting on bed of kelp (Laminariales), Prince William Sound, Alaska, USA, Pacific Ocean
Pair of Couch's spadefoot toads (Scaphiopus couchii) mating in water, Texas, USA. June
Red tailed hawk (Buteo jamaicensis) juvenile female, head portrait, Texas, USA. April
Galapagos sea lion (Zalophus wollebaeki) female with pup lying on rocks, Espanola Island, Galapagos, South America
Snowy owl (Bubo scandiaca) female, head portrait, Canada. January
Snowy owl (Bubo scandiaca) female, in flight in winter, Canada. January
Arctic fox (Vulpes lagopus), female blue colour morph in winter coat, during snow blizzard. Baited with small amount of natural food (dried fish). Hornstrandir, Iceland. February
Red fox (Vulpes vulpes) vixen walking through snow, Derbyshire, UK. January
Female Bearded reedling (Panurus biarmicus) perched on frost-covered reeds, Finland. February
A female kingfisher (Alcedo atthis) perched on a branch in snow. Leeds, Yorkshire, UK. January
Magnificent bird of paradise (Cicinnurus magnificus) male, leaning back and presenting his breast shield and raised cape to two females, courtship display, Arfak Mountains, West Papua, Indonesia
Pair of Wilson's birds of paradise (Cicinnurus respublica), male performing courtship display to female from a small sapling in the middle of his display court, Batanta Island, West Papua
Lioness (Panthera leo) lying in long grass on the savannah, portrait, Okavango Delta, Botswana, Africa
Female Gentoo penguin (Pygoscelis papua) with chick on the nest, Port Lockroy, Goudier Island, Antarctic Peninsula, Antarctica
African wild dog (Lycaon pictus) alpha female, interacting with two of her pups, aged 4 weeks, Okavango Delta, Northern Botswana, Africa
African wild dog (Lycaon pictus) alpha female suckling her 12 pups, aged 4 weeks, Okavango Delta, Northern Botswana, Africa
Pair of Lava gulls (Leucophaeus fuliginosus) during courtship, Genovesa Island, Galapagos Islands, Ecuador
Puma (Puma concolor) female, grooming her cub, Torres del Paine National Park, Magallanes, Chile
Leatherback turtle (Dermochelys coriacea) female, in shallow river, trying to find her way back to the sea after laying eggs on a sandbar, Grande Riviere, Trinidad Island, Trinidad & Tobago, Caribbean
Puma (Puma concolor) female, walking in front of Torres del Paine massif, Torres del Paine National Park, Patagonia, Chile
Puma (Puma concolor) female, running in deep fresh, snow, Torres del Paine National Park, Patagonia, Chile
Puma (Puma concolor) female with two cubs, aged six months, sitting in fresh snow, Torres del Paine National Park, Patagonia, Chile
Puma (Puma concolor) female, walking in deep fresh, snow, Torres del Paine National Park, Patagonia, Chile
Young male Lion and lioness (Panthera leo) resting in tree during middle of the day to escape the heat. Acacia woodland near Ndutu, Ngorongoro Conservation Area / Serengeti National Park border
Female leopard (Panthera pardus) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa
Arctic fox (Vulpes lagopus), female blue colour morph in winter coat, walking across snow covered landscape. Baited with small amount of natural food (dried fish). Hornstrandir, Iceland. February
Grey-headed flying-fox bat (Pteropus poliocephalus), female, hanging the other way to defecate or urinate, Yarra Bend Park, Victoria, Australia. Cropped
Female Eastern bluebird (Sialia sialis) spreading wings for balance after landing on a fruiting Holly (Ilex sp)Female Eastern bluebird (Sialia sialis) spreading wings for balance after landing on a fruiting Holly (Ilex sp.) bush, Freeville, New York, USA. January
Female Ostrich (Struthio camelus) feeding on desert plants, Kgalagadi Transfrontier Park, South Africa
Two Lionesses (Panthera leo) nuzzling surrounded by three cubs, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Female Bison (Bison bison) with young calf, walking over snow in front of frost-covered tree, Yellowstone National Park, USA. February
Giant panda (Ailuropoda melanoleuca) cub Yuandudu, aged 8 months, portrait, Beauval ZooPark, France, April, 2022. Captive
Female Nursery web spider (Pisaura mirabilis) carrying egg sac, Peak District National Park, Derbyshire, UK. June
Female Praying mantis (Mantis religiosa) crawling on thistle, Lorraine, France. August
Female Leopard (Panthera pardus) carrying cub in her mouth to their new den, Kgalagadi Transfrontier Park, South Africa
Female Cheetah (Acinonyx jubatus) with juveniles crossing grassland in rain storm, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Female Olive ridley turtle (Lepidochelys olivacea) coming ashore at dusk to lay eggs, Oaxaca, Mexico, Pacific Ocean
Female Olive ridley turtles (Lepidochelys olivacea) coming ashore at night in large numbers to lay eggs during arribada mass nesting, Oaxaca, Mexico
Gray whale (Eschrichtius robustus), mother and calf, swimming at surface, Magdalena Bay, Baja California, Mexico, Pacific Ocean
RF - Female Scalloped hammerhead shark (Sphyrna lewini) surrounded by Pacific creolefish (Paranthias colonus), Wolf Island, Galapagos National Park, Galapagos Islands, Pacific Ocean
Pair of Peppermint basslets (Liopropoma rubre) spawning at sunset, with eggs floating alongside the female, Grand Cayman, Cayman Islands, Caribbean Sea
Roe deer (Capreolus capreolus) doe with fawn standing in hedgerow, Fife, Scotland, UK. September
Female polar bear (Ursus maritimus) with two cubs walking in a line across snow, Churchill, Canada. November
Female Kingfisher (Alcedo atthis) covering chicks, aged one day, with her wings in an artificial nest, Italy
Three female Putorana snow sheep (Ovis nivicola borealis) climbing up rocky mountainside with lamb following, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Gravid female Timber rattlesnakes (Crotalus horridus) basking to bring young to term, Pennsylvania, USA
Female Serval (Leptailurus serval) portrait. Masai Mara national reserve, Kenya, East Africa
White-tailed deer (Odocoileus virginianus) doe with fawn in woodland, Acadia National Park, Maine, USA. July
