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Female Grizzly bear (Ursus arctos horribilis) with cub among wildflowers, Grand Teton National Park, Wyoming, USA. June
Female Black woodpecker (Dryocopus martius), pecking at tree trunk, Finland. September
Young female Grey wolf (Canis lupus) in meadow, Finland. July
Female Silver-washed fritillary butterfly (Argynnis paphia) nectaring on Wild angelica (Angelica sylvestris), Finland. July
Giant panda (Ailuropoda melanoleuca) cubs Yuandudu and Huanlili, aged 8 months, playing together, Beauval ZooPark, France, April, 2022. Captive
Giant panda (Ailuropoda melanoleuca) cubs, Yuandudu and Huanlili, aged 8 months, side by side, Beauval ZooPark, France, April, 2022. Captive
Red fox (Vulpes vulpes) vixen standing in fork of tree on snowy night, Vertes Mountains, Hungary
Red fox (Vulpes vulpes) vixen looking in through window of traditional cottage house, Vertes Mountains, Hungary
Female Red fox (Vulpes vulpes) cautiously staring at the camera from the base of a moss-covered tree, Hungary. January
Female Red fox (Vulpes vulpes) walking along tree trunk in heavy fog at night. Light source behind the subject creates dramatic, volumetric lighting, Hungary, January
Female Red fox (Vulpes vulpes) waiting to be fed at the back entrance of a house, Hungary. January
Female Red fox (Vulpes vulpes) observing the yard from the top of the old stone wall, trying to determine whether it is safe to enter, Hungary. January
Female Abyssinian ground hornbill (Bucorvus Abyssinicus) walking along riverbank, Allahein River, The Gambia
Female Goldenrod crab spider (Misumena vatia) camouflaged on Yellow flag iris (Iris pseudacorus) flower waiting for prey to land, Kenfig National Nature Reserve, Glamorgan, Wales, UK, June
Three Lined seahorses (Hippocampus erectus), two male and one female, feeding on plankton in an alkaline saltwater pond, The Bahamas, Caribbean
A female kingfisher (Alcedo atthis) fishing / diving into an ice hole in winter. Leeds, Yorkshire, UK. January
Lowland gorilla (Gorilla gorilla) mother and young in forest, Loango National Park, Gabon. January
Baltimore oriole (Icterus galbula) female perched on Berlandiers fiddlewood (Citharexylum berlandieri). South Padre Island, Texas, USA. April
Ruby-throated hummingbird (Archilochus colubris) female nectaring on Maximilian sunflower (Helianthus maximilianii). Hill Country, Texas, USA
RF - Broad-bodied chaser dragonfly (Libellula depressa) resting on reeds. Cornwall, England, UK. May. (This image may be licensed either as rights managed or royalty free.)
Beautiful demoiselle damselfly (Calopteryx virgo) female resting on Yellow iris (Iris pseudacorus) bud. Cornwall, England, UK. May
Jaguar (Panthera onca) female stealthily entering the river as she begins to hunt caiman. Northern Pantanal Cuiaba River, Mato Grosso, Brazil
RF - Cheetah (Acinonyx jubatus) female patrolling territory. Long Gully, Ngorongoro Conservation Area, Tanzania. April
Leopard (Panthera pardus) female drinking at night. South Luangwa National Park, Zambia
Southern Tamandua (Tamandua tetradactyla) female carrying young on her back, Northern Pantanal, Mato Grosso State, Brazil, South America
Adult Fossa (Crytoprocta ferox) female portrait. Kirindy Forest, western Madagascar, November
Dominant male Chimpanzee (Pan troglodytes troglodytes) walking in the mangrove, with female carrying her infant, Conkouati-Douli National Park, Republic of Congo, Africa
Female Chimpanzee (Pan troglodytes troglodytes) climbing in trees carrying her infant, Conkouati-Douli National Park, Republic of Congo, Africa
Lisbeths brooding anemone (Epiactis lisbethae), young clinging on to their mother, Vancouver Island, British Columbia, Canada, Pacific Ocean
Female Eclectus parrot (Eclectus roratus) feeding in rainforest, Cape York, Queensland, Australia
Female Plum-headed finch (Neochima modesta) feeding in long grass, Dalby, Queensland, Australia
Female Yellow-footed rock wallaby (Petrogale xanthopus) fending off advances from male (right), Idalia National Park, Queensland, Australia
Dominant male Chimpanzee (Pan troglodytes troglodytes) in the mangrove, with female and infant vocalising, Conkouati-Douli National Park, Republic of Congo, Africa
Female Galapagos sea lion (Zalophus wollebaeki), sleeping together, likely to be related, Santa Fe Island, Galapagos, South America
Galapagos sea lion (Zalophus wollebaeki) mother with newborn pup sitting on beach, Espanola Island, Galapagos, South America
Female California sheephead (Semicossyphus pulcher) and miscellaneous reef fish swimming in a forest of Giant kelp (Macrocystis pyrifera) off Santa Barbara Island, California, USA
Female European hoverfly (Helophilus pendulus) pollinating Tansy (Tanacetum vulgare) in flower, Belgium. August
Female Fox (Vulpes vulpes) caught in a shaft of light on the porch of a farmhouse, Verteskozma, Hungary
Female Fox (Vulpes vulpes) walking on tree trunk in heavy snowstorm in the dark, Verteskozma, Hungary
Female Red fox (Vulpes vulpes) foraging inside a rotting tree trunk, Verteskozma, Hungary. Hungary Nature Photographer of the Year Competition 2021 Highly Commended - Mammals Behaviour category
Female Cuviers gazelle (Gazella cuvieri) calf, age 6 months, with its mother, Djebel Ouarkziz, Southern Morocco, Africa. Customized camera-trap
Fishing cat (Prionailurus viverrinus) with two kittens, age 4 weeks, learning to hunt for fish in wetlands, photographed during a release project to relocate Fishing cats affected by habitat loss
A female kingfisher (Alcedo atthis) fishing / diving into an ice hole in winter. Leeds, Yorkshire, UK. January. Sequence 1 of 4
A female kingfisher (Alcedo atthis) fishing, flying out of an ice hole in winter. Leeds, Yorkshire, UK. January
Pair of Mute swans (Cygnus olor) at sunrise on misty lake. Poland, March
Minors chameleon (Furcifer minor), female, climbing down dried plant stalk, Madagascar
Scaly breasted lorikeet (Trichoglossus chlorolepidotus) pair allopreening in branch of tree, Toowoomba, Queensland, Australia
Wild boar (Sus scrofa) sow with piglets. Forest of Dean, Gloucestershire, England, UK. March