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White-backed woodpecker (Dendrocopos leucotos), female excavating nest in birch tree, expelling wood chips from hole, Finland, April
Sheephead (Semicossyphus pulcher) female, and Garibaldi, (Hypsypops rubicundus) in forest of giant kelp (Macrocystis pyrifera) off Santa Barbara Island, California, USA
Iberian lynx (Lynx pardinus) female with playful cub, Sierra de Andujar Natural Park, Andalucia, Spain
Tigress (Panthera tigris tigris) with her four cubs from Chandrapur district, Maharashtra, India. She lives in an area known for human-animal conflict
Small emerald damselfly (Lestes virens) female resting, dew droplets on body, close up. Klein Schietveld, Brasschaat, Belgium. August
Roe deer (Capreolus capreolus) doe sitting amongst logs. Peerdsbos, Brasschaat, Belgium. July
Roe deer (Capreolus capreolus) doe standing in grassland. Peerdsbos, Brasschaat, Belgium. July
Silky sifaka (Propithecus candidus) female with baby sitting amongst branches in understorey of mid-altitude montane rainforest. Marojejy National Park, north east Madagascar
A split level photo of a female southern stingray (Dasyatis americana) swimming over seabed. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Red deer (Cervus elaphus) stag bellowing during rut in grassland, doe in background
Bumblebee (Bombus sp) queen feeding on Crocus (Crocus korolkowii), covered in pollen. Ansob Pass, Tajikistan. April
Ivy bee (Colletes hederae) female collecting pollen from Ivy (Hedera helix). Surrey, England, UK. October
Wool carder bee (Anthidium manicatum) female entering Rusty foxglove (Digitalis ferruginea) flower. Pollen transferring to head from stamens on upper lip of flower
Buff tailed bumblebee (Bombus terrestris) queen nectaring on Rhododendron (Rhododendron sp), pollen from anthers on backside and legs. Surrey, England, UK. March
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
Crowned lemur (Eulemur coronatus) two females on ground in forest, Ankarana NP, Madgascar
Chinese white cheeked gibbon (Nomascus leucogenys) female hanging from tree, carrying baby aged two weeks. Parents released into wild from captive breeding programme
Common seahorse (Hippocampus kuda) female wrapping her prehensile tail around piece of wood on the seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
RF- American robin (Turdus migratorius) female feeding in Black hawthorn (Crataegus douglasii). Grand Teton National Park, Wyoming, USA
Snow monkey or Japanese macaque (Macaca fuscata) female with baby clinging to her back. Jigokudani, Japan, February
Red deer (Cervus elaphus) herd of female does and young crossing water, Jura, Scotland, UK, September
Red deer (Cervus elaphus) female does and young crossing water, Jura, Scotland, UK, September
Kingfisher (Alcedo atthis) female emerging from water with fish, Hampstead Heath, London, England, UK, September
Red deer (Cervus elaphus) stag bellowing, surrounded by hinds, Surrey, UK, September
Female Chinese water deer (Hydropotes inermis) Cley, Norfolk, UK, August
Roe deer (Capreolus capreolus) doe in Scots pine woodland, Cairngorms National Park, Scotland, UK, August
Female Meadow brown butterfly (Maniola jurtina) feeding on thistle, Lewisham, London, UK, August
Sperm whales (Physeter macrocephalus) aerial shot showing much larger male on right, with female nursery group. Baja California, Mexico
Red leaf monkey (Presbytis rubicunda) female feeding in Strangler fig tree (Ficus dubia) Gunung Palung National Park, Kalimantan, Borneo, Indonesia
Coquerels sifaka (Propithecus coquereli) female with young on back climbing in bamboo, Anjajavy Private Reserve, north west Madagascar
Emerald damselfly (Lestes sponsa), female resting, Broxwater, Cornwall, UK. August
Pronghorn antelope (Antilocapra americana) females, Saskatchewan, Canada
Harp seal (Phoca groenlandica), adult female on the ice, Magdalen Islands, Canada
Silver washed fritillary butterfly (Argynnis paphia) female in flight with thistles, Uusimaa, Finland, August
Blackbuck (Antelope cervicapra), wide angle ground perspective of female. Camera trap image. Tal Chhapar Wildlife Sanctuary, Rajasthan, India
Jaguar (Panthera onca) female portrait, native to Southern and Central America, captive
Leaf-cutting bee (Megachile species) carrying leaf section. Surrey, England, UK, July
Grey seal (Halichoerus grypus) large female exhales a stream of bubbles as she sleeps on shallow seaweeds (Fucus serratus) Lundy Island, Devon, UK, Bristol Channel, August
Leopard (Panthera pardus) female grooming cub age 6 months Little Kwara, Botswana June
Impala (Aepyceros melampus) leaping, Little Kwara, Botswana June
Leatherback turtle (Dermochelys coriacea) female returning to sea after laying eggs, Trinidad and Tobago, April
Common reed bunting (Emberiza schoeniclus) attacking Common cuckoo (Cuculus canorus), female, Braunschweig, Lower Saxony, Germany, May
Snowy owl (Bubo scandiaca) female face close up, Canada February
Raft Spider (Dolomedes fimbriatus) female resting on the surface of a moorland pool, photographed with a fisheye lens to show surrounding habitat. Nordtirol, Austrian Alps, June
Pride of Lions (Panthera leo) on termite mound at sunset, Okavango Delta, Botswana
Mother with young calf Sperm whale (Physeter macrocephalus) Dominica, Caribbean Sea, Atlantic Ocean
Large copper butterfly (Lycaena dispar), female, Finland, July
Meadow Brown Butterfly (Maniola jurtina) female basking, wings open, Southwest London, UK, July