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Aerial view of Lesser flamingos (Phoeniconaias minor), mating display Lake Magadi, Rift Valley, Kenya
Andean Cock-of-the-rock male (Rupicola peruvianus) at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
African grey hornbill (Lophoceros nasutus) calling in display, perched on tree snag. Janjanbureh, Gambia
Waved albatrosses (Phoebastria irrorata) courting, Punta Suarez, Espanola Island, Galapagos
California least terns (Sternula antillarum browni) courtship behaviour, Bolsa Chica Ecological Reserve, California, USA June
Common Merganser (Mergus merganser) female with ducklings, in threat posture at the approach of a second female, Lansing, New York, USA
Leopard (Panthera pardus) pair mating behaviour, Okavango Delta, Botswana
Black Grouse (Lyrurus tetrix) fighting on icem Finland. March
Pair of courting Egg-eating sea snakes / Turtleheaded sea snakes (Emydocephalus annulatus) New Caledonia, Pacific Ocean
Eurasian rollers (Coracias garrulus) courting on branch, Hungary. June
King cormorants (Phalacrocorax atriceps) in courtship behaviour at their nest site. Saunders Island, Falkland Islands
Red deer (Cervus elaphus) stag bellowing during rut in grassland, doe in background
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
Amur / Siberian tiger (Panthera tigris altaica) pair in courtship, in snow. Captive in tiger park, Heilongjiang Province, China. January
Waved albatross (Phoebastria irrorata) pair in courtship display at nest site, Punta Suarez, Espanola Island, Galapagos
Black-browed albatross pair (Thalassarche melanophris) mutual grooming during copulation. New Island, Falkland Islands, November
Nurse sharks (Ginglymostoma cirratum) three in a courtship dance at sunrise in a mangrove area near Eleuthera, Bahamas
Magellanic penguin (Spheniscus magellanicus) calling at the entrance of burrow, courtship display, Carcass Island, Falkland Islands, October
Laysan albatross (Phoebastria immutabilis), courtship ritual, Eastern island, Midway Atoll National Wildlife Refuge, Hawaii
Brown Hare (Lepus europaeus), Bayern, Germany. April
Green mandarinfish (Synchiropus splendidus) pair courting and mating at sunset, Sulu Sea, Philippines
American elk (Cervus elaphus canadensis) stag sticking tongue out, Yellowstone National Park, Wyoming, USA
Sperm whale (Physeter macrocephalus) young male trying to copulate with a female. The calf of the female was in the way so the male is trying to chase it away. Dominica
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Andean Cock-of-the-rock male {Rupicola peruvianus} at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Long-tailed manakin (Chiroxiphia linearis) perched on lek (display perch), Costa Rica
Sooty Albatross (Phoebetria fusca) group courting on high inland ridge. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic. Endangered
RF - Red deer (Cervus elaphus) stags fighting, Holkham Park, Norfolk, England, UK, September
Black Grouse (Tetrao tetrix) lek with male displaying and females around in winter, Tver, Russia. April
Black Grouse (Tetrao tetrix) males fighting in winter, Tver, Russia. April
Horned puffins (Fratercula corniculata) pair interacting by touching bills while perched on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Black grouse (Tetrao tetrix) male displaying at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (tetrao tetrix) males fighting at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (Tetrao tetrix) males displaying at a lek site at dawn, Cairngorms National Park, Scotland, UK, April
Black necked grebe (Podiceps nigricollis) pair at night, performing their courtship dance during the mating season, with street lights reflected in the water. The Netherlands.April 2014
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Brown hare (Lepus europaeus) two males chasing female in snow covered field during a blizzard. Derbyshire, England, UK, February. Winner of British Seasons category of BWPA competition 2014
Moorland Clouded Yellow butterfly (Colias palaeno) male and female, Joutsa (formerly Leivonmaki), Finland, July
Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April
RF- Black Grouse (Lyrurus tetrix) displaying. Finland. March
Common cranes (Grus grus) displaying, Hula Valley, Northern Israel, January
Andean cock of the rock (Rupicola peruviana) male in courtship ritual, Andean Mountains, Merida, Venezuela
Wolf Spiders (Pardosa sp. ), male RIGHT waving palps in courtship displayWolf Spiders (Pardosa sp.), male RIGHT waving palps in courtship display. Peak District National Park, Derbyhsire, UK. May
Northern gannets (Morus / Sula bassanus) portrait of individual with a courting pair in foreground, UK, Bass Rock, Scotland
