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Lesser kestrel (Falco naumanni) male flying past during courtship with Mole cricket in its talons. Extremadura, Spain
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February
Red deer (Cervus elaphus) stag bellowing during rut, Derbyshire, UK
Greater sage-grouse (Centrocercus urophasianus) male displaying on a lek in snow, Sublette County, Wyoming, USA. April
Great crested grebe (Podiceps cristatus) pair, courtship dance at dawn, Sierra de Grazalema Natural Park, Andalusia, Spain. April
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Iberian ibex (Capra pyrenaica), pair with male top right, during rutting season. Sierra Nevada National Park, Andalusia, Spain. October
Pair of Waved albatrosses (Phoebastria irrorata) billing as greeting. Punta Suarez, Espanola Island, Galapagos Islands, Ecuador
Wine throated hummingbird (Selasphorus ellioti) male, courtship display, Cloud forest, Atitlan Volcano, Solola, Guatemala
Ruddy duck (Oxyura jamaicensis) male in courtship display. Valkenhorst nature reserve, Valkenswaard, The Netherlands. May
Humpback whales (Megaptera novaeangliae) pair engaging in courtship behavior, dancing for hours, off Tubuai, French Polynesia, Pacific Ocean
Bluethroat (Luscinia svecica cyanecula) male, singing as a courtship display, Germany
Magnificent frigatebirds (Fregata magnificens) pair in courtship display, male with gular sac inflated, North Seymour Island, Galapagos National Park, Galapagos Islands
Club-winged manakin (Machaeropterus deliciosus) male, courtship display at lek, Choco rainforest, Pichincha, Ecuador
European Common adder (Vipera berus) pair courting and retreating under gorse (Ulex sp). Buckinghamshire, England, UK. May
Pair of Red mason bees (Osmia bicornis) mating on wallflower in garden. Hertfordshire, England, UK. May. Focus Stacked
Brown hare, (Lepus europaeus), male and female displaying courting behaviour, Islay, Scotland, UK. March
Brown hare, (Lepus europaeus), males in pursuit of female that is in season, Islay, Scotland, UK. March
Pair of Spotted unicornfish (Naso brevirostris) courtship display above the reef, Ari Atoll, Maldives, Indian Ocean
Two Ruffs (Philomachus pugnax) male, competing at a lek, Vardo, Norway. May
Vermilion flycatcher (Pyrocephalus obscurus) male, courtship display, Texas, USA. May
American flamingo (Phoenicopterus ruber) performing courtship wing flashing, Quinta Playa, Isabela Island, Galapagos Islands, Ecuador
Magnificent bird of paradise (Cicinnurus magnificus) male, leaning back and presenting his breast shield and raised cape to two females, courtship display, Arfak Mountains, West Papua, Indonesia
Wilson's bird of paradise (Cicinnurus respublica) male, courtship display, Batanta Island, West Papua, Indonesia
Pair of Wilson's birds of paradise (Cicinnurus respublica), male performing courtship display to female from a small sapling in the middle of his display court, Batanta Island, West Papua
Vogelkop bowerbird (Amblyornis inornatus) male, at his bower surrounded by orange leaves and petals and other objects he has collected, Arfak Mountains, Vogelkop Peninsula, West Papua
Wandering albatross (Diomedea exulans) male, standing with wings spread in courtship display, Prion Island, South Georgia
Magnificent riflebird (Ptiloris magnificus alberti) male, courtship display, Piccaninny Plains Sanctuary, Cape York Peninsula, Queensland, Australia
Pair of Lava gulls (Leucophaeus fuliginosus) during courtship, Genovesa Island, Galapagos Islands, Ecuador
Flock of American flamingos (Phoenicopterus ruber) performing courtship strutting and wing flashing, Quinta Playa, Isabela Island, Galapagos Islands, Ecuador
Male Beautiful sunbird (Cinnyris Pulchellus) courtship display, Allahein River, The Gambia
Two male Black grouse (Tetrao tetrix) fighting for territory on lek, Cairngorms National Park, Scotland, UK. May
Male Black grouse (Tetrao tetrix) performing courtship display at lek, Cairngorms National Park, Scotland, UK. April
Male Western capercaillie (Tetrao urogallus) calling during lek mating, Finland. March
Pair of adult Red-necked grebe (Podiceps grisegena) calling, Finland. May
Close up of male Western capercaillie (Tetrao urogallus) during lek mating, Finland. March
Pair of Great crested grebes (Podiceps cristatus) performing the weed dance during courtship, The Netherlands, Europe. March
Female Yellow-footed rock wallaby (Petrogale xanthopus) fending off advances from male (right), Idalia National Park, Queensland, Australia
Red-throated diver (Gavia stellata) pair displaying on the water, Vaala, Finland. May
Pair of Waved albatross (Phoebastria irrorata) courtship, Espanola Island, Galapagos, South America
Silvery grebe (Podiceps occipitalis) attempting to join Hooded grebes (Podiceps gallardoi) during a breeding display, Patagonia National Park, Santa Cruz, Argentina. Cropped
White stork (Ciconia ciconia) pair displaying on their nest built on the chimneys of Knepp Castle, Sussex, UK, June
Gentoo penguins (Pygoscelis papua) pair, Sea Lion Island, Falkland Islands
Black-browed albatross (Thalassarche melanophris) pair in courtship. Falkland Islands
Waved albatross (Phoebastria irrorata), pair mutual preening in courtship, Espanola Island, Galapagos Islands
Kingfisher (Alcedo atthis) male passing fish to female, courtship behaviour, Lorraine, France, March
Bilingual frog (Crinia bilingua) adult male calling, Adelaide River, Northern Territory, Australia, January
Two Black-browed albatross (Thalassarche melanophris) rub their bills together - part of a bonding ritual. West Point Island, The Falkland Islands
