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Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
Annas hummingbird (Calypte anna) male dancing in flight and displaying his bright pink throat feathers to a female, Montana, USA. June
Firefly (Lamprohiza splendidula) light trails of males in forest at dusk, Bavaria, Germany. July
Greater Prairie-Chicken (Tympanuchus cupido) male displaying on lek. Fort Pierre National Grassland, South Dakota, USA. When displaying
Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Red fox (Vulpes vulpes) pair nuzzling. London, UK. February
Male Black grouse (Tetrao / Lyrurus tetrix) in flight. Kuusamo, Finland, April
Boat-tailed grackle pair (Quiscalus major) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Lawess parotia (Parotia lawesii) male displaying. Papua New Guinea
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
Red Deer (Cervus elaphus) stags fighting during the rutting season. Bushy Park, London, UK. October
Manchurian crane (Grus japonensis) pair in courtship dance during snowstorm. Hokkaido, Japan. March
Fireflies (Luciola parvula himebotaru) flashing at night for courtship and reproduction. Gifu, Japan. Composite image
Eared grebes (Podiceps nigricollis) stand erect in water during courtship, North Park, Colorado, USA. June
Amur / Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snowAmur/ Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snow. Captive in tiger park, Heilongjiang Province, China. February
Fallow deer (Dama dama) buck bellowing in autumn vegetation, Bradgate Park, Leicestershire, UK, October
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Sarus crane (Grus antigone), male displaying, Keoladeo NP, Bharatpur, India
Lioness (Panthera leo), aggressively rebuffing male lion banks of the Luangwa river, South Luangwa National Park, Zambia
Northern gannets (Morus bassanus) performing sky-pointing display with outspread wings as part of courtship, Cape St. Marys Ecological Reserve, Newfoundland, Canada. June
Red-breasted Mergansers (Mergus serrator) male in foreground performing courtship display to female in background (left), Aurora, New York, USA, April
Western Grebe (Aechmorphus occidentalis), pair during rushing courtship display in which they run across waters surface in synchrony, near Escondido, California, USA, January
American oystercatchers (Haematopus palliatus) courting pair performing Piping Display, calling as they walk along together, Fort DeSoto Park, Florida, USA, March
White stork (Ciconia ciconia) pair performing an up-down display with bill clattering on their nest in an Oak tree, Knepp estate, Sussex, UK, April 2019
Red deer (Cervus elaphus) herd during rut in morning light. Derbyshire, England, UK. October
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Jaguars (Panthera onca) courting pair on a sand bank. Cuiaba River, Northern Pantanal, Mato Grosso, Brazil
Two rainbow lorikeets (Trichoglossus moluccanus) on branch, Werribee, Victoria, Australia
Montezuma oropendola (Psarocolius montezuma) hanging upside down as part of a courtship display. Costa Rica
Kori bustard (Ardeotis kori) male courtship display, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Clarion Lizard (Urosaurus clarionensis) displaying, IUCN Vulnerable, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Hoopoe (Upupa epops) male feeding female in courtship display, Pusztaszer, Hungary, April
Sperm whale (Physeter macrocephalus) young male trying to copulate with a female. The calf of the female was in the way so the male is trying to chase it away. Dominica
Andean Cock-of-the-rock male {Rupicola peruvianus} at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Western capercaillie (Tetrao urogallus) male calling at lek, with female, Tver, Russia. May
Black Grouse (Tetrao tetrix) males fighting at lek in the snow, Tver, Russia. April
Gunnison sage-grouse (Centrocercus minimus) male displaying at a lek. Gunnison County, Colorado. April
Royal terns (Sterna maxima) pair crossing bills during courtship behaviour, Fort DeSoto Park, Florida, USA
Whooper swan (Cygnus cygnus) pair on loch in heart shape, Cairngorms National Park, Scotland, UK, November
Fallow deer (Dama dama) buck bellowing at dawn during the rut, Cheshire, UK, October
Satellite male Ruff (Philomachus pugnax) in full display at the lek. Varanger, Finmark, Norway, May
Lesser kestrel (Falco naumanni) male flying past during courtship with Mole cricket in its talons. Extremadura, Spain
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February