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Chocolate working cocker spaniel in snow, Wiltshire, UK
Barn owl (Tyto alba) in flight, hunting, Hampshire, England, UK. Captive
Snowy owl (Bubo scandiaca) female, with lights behind, Canada, February
Golden Eagle (Aquila chrysaetos) juvenile in flight, Norway, November
Wandering albatross (Diomedea exulans) in flight over the ocean, Drake Passage, Southern Ocean
Blue whale (Balaenoptera musculus) diving beneath ocean surface. Indian Ocean, Sri Lanka
Atlantic puffin (Fratercula arctica) close up of head, Fair Isle, Shetland, Scotland, UK, July
Atlantic Puffins (Fratercula arctica) bill rubbing, Skomer Island, Wales, UK, April
RF - Lion (Panthera leo) head portrait at night, Zimanga private game reserve, KwaZulu-Natal, South Africa. (This image may be licensed either as rights managed or royalty free.)
Eurasian flamingos (Phoenicopterus roseus) lined up on Pulicat Lake, Tamil Nadu, India, January 2013
Common kestrel (Falco tinnunculus) perched on a branch, Valencia, Spain, February
Golden working cocker spaniel puppy sitting amongst bluebells in beech woodland. Micheldever Woods, Hampshire, UK
Male Saiga antelope (Saiga tatarica) in winter, The Black Lands National Park, Kalmykia, Russia
European roller (Coracias garrulus) in flight, Pusztaszer, Hungary, May
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Black working cocker spaniel in bluebells in beech woodland, Micheldever Woods, Hampshire, UK
Angular roughshark (Oxynotus centrina) a deepsea species living at 80-300m depth, caught by fishermen and released, Costa Brava, Catalunya, Spain, Mediterranean Sea
Deep sea jellyfish (Periphylla periphylla) juvenile, Trondheimsfjord, Norway, Atlantic ocean
Great Spotted Woodpecker in snow (Dendrocopos major), Scotland, February
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Whooper swans (Cygnus cygnus) three coming into land, Hokkaido, Japan, February
Oystercatcher (Haematopus ostralegus) by the coast, Island of Runde, Norway July
Black-throated Diver (Gavia arctica) Rokua, Finland, May
Japanese cranes (Grus japonensis) displaying in snow, Hokkiado, Japan, February
Willow Grouse (Lagopus lagopus) male perched on snow, Finland, March. Bookplate from Danny Greens The Long Journey North
Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016
RF- Goldfinch (Carduelis carduelis) reflected in pool, Worcestershire. November
Portrait of a Green sea turtle (Chelonia mydas) in morning light. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea
Tropical shield mantis (Choeradodis sp. ) (Possibly C. rhombicollis) Canande, EsmeraldasTropical shield mantis (Choeradodis sp.) (Possibly C. rhombicollis) Canande, Esmeraldas, Ecuador
Great tit (Parus major) perched on branch, ready to fly, Moselle, France, February
Arctic tern (Sterna paradisaea) hovering above the ocean, Svalbard, Norway June
RF - Red Squirrel (Sciurus vulgaris) on log in snow. Scotland, UK. December
Golden eagle (Aquila chrysaetos) sub-adult flying, Strathdearn, Inverness-shire, Scotland, UK, August
Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Pine marten (Martes martes) portrait, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Portrait category of the BWPA (British Wildlife Photography Awards) Competition 2016
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
Dalmatian Pelican (Pelecanus crispus) portrait, close-up of eyes, Lake Kerkini, Greece. February
Herb robert (Geranium robertianum) flower, Coombe Valley, Cornwall, UK, May
Goldfinch (Carduelis carduelis) feeding on teasel in falling snow, Leicestershire, UK, February
Eyrean Grasswren (Amytornis goyderi) in typical hopping motion, Andado Station, Northern Territory, Australia, November
Grey heron (Ardea cinerea) wading at night, reflected in water. Lincolnshire, UK, September
Wren (Troglodytes troglodytes) singing, early morning in spring, Broxwater, Cornwall, UK. April
Robin (Erithacus rubecula) feeding on Gelder Rose berries, Germany. January, January
Wren (Troglodytes troglodytes) on frost covered fern, Dunham Massey, Cheshire, UK. Winner of the Habitat Category of the British Wildlife Photographer of the Year Awards (BWPA) 2017
Portrait of Sally lightfoot crab (Grapsus grapsus) on a beach. San Cristobal Island, Galapagos Islands, Ecuador. Tropical East Pacific Ocean
Caspian Tern (Sterna caspia) in flight, diving for prey. Lake Ellesmere, Canterbury, South Island, New Zealand
Common toadflax (Linaria vulgaris), orange nectar guides on lower lip and long spur containing nectar. Surrey, England, UK. Focus stacked
Ptarmigan (Lagopus muta) male on snowy ridge on Cairngorm mountain, Cairngorms National Park, Scotland, UK, March
Tree sparrow (Passer montanus) feeding a fledgling on a branch of Hawthorn (Crataegus monogyna) blossom, Perthshire, Scotland. June 2013
Raven (Corvus corax) calling in the snow, Kemijarvi, Finland, February
Sword-billed hummingbird (Ensifera ensifera) hovering in flight, North-Ecuador
Mountain hare (Lepus timidus) adult resting in snow hole, Scotland, UK, February
Golden working cocker spaniel puppy amongst bluebells in beech woodland, Micheldever Woods, Hampshire, UK
Cow killer velvet ant (Dasymutilla occidentalis) on white background, Tuscaloosa County, Alabama, USA September
Red fox (Vulpes vulpes) rear view sitting on grass close to its den, Derbyshire, UK
Female Black-footed cat (Felis nigripes) in the desert at night, Karoo, South Africa. Taken on location for the BBC series Big Cats
Great crested grebe (Podiceps cristatus) adult with young on the back in the first morning light Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. May
Wood pigeon (Columba palumbus) flying with nesting material in beak, Lorraine, France, April
Arctic fox (Vulpes lagopus), blue colour morph in winter coat walking through snow. Hornstrandir Nature Reserve, Iceland. February
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
Deepsea jellyfish (Periphylla periphylla) juvenile, , Trondheims fjord, Norway, Atlantic ocean
Rabbit (Oryctolagus cunniculus) juvenile grooming ears, Wiltshire, UK, May. Sequence 1 of 3
Pallass cat (Otocolobus manul) licking lips whilst running over ice. East Mongolia. February
Sparrowhawk (Accipiter nisus) adult female, Norway, October
Bush cricket / Katydid (Tettigoniidae ), female leaf mimic ovipositing into branch. Manu Biosphere Reserve, Peru
Short-finned pilot whale (Globicephala macrorhynchus) calf with fetal lines characteristic of newborn specimens, with pod. Tenerife, Canary Islands
Arctic fox (Alopex lagopus) camouflaged in winter pelage. Svalbard, Norway, April
Flying fish (Exocoetidae) in Sargasso Sea, Atlantic Ocean
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Geraldton wax flower (Chamelaucium uncinatum), nectar fluorescing in UV light. Western Australia. Controlled conditions, focus stacked. Series 1/2
Flame lily (Gloriosa superba), timelapse sequence from opening bud to flowering, tepals becoming reflexed. Flower pollinated by butterflies. Controlled conditions
Hummingbird hawk-moth (Macroglossum stellatarum) nectaring on Loofah (Luffa sp) flower. Xinjiang, China
Chinese sturgeon (Acipenser sinensis), captive in aquarium. Adults are no longer able to swim upriver to spawn due to Gezhouba and Three Gorges dams
Bubble eye goldfish (Carassius auratus) with upward pointing eyes and two large fluid-filled sacs.. In captivity, Shanghai Zoo, Shanghai, China
Bruising webcap mushroom (Cortinarius purpurascens) Annagarriff Wood NNR, Peatlands Park, County Armagh, Northern Ireland. November
Rock Ptarmigan (Lagopus muta) portrait, Inari, Finland, September
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Violet-tailed sylph hummingbird (Aglaiocercus coelestis) in flight, about to land. Ecuador, May
Pelagic thresher shark (Alopias pelagicus) being cleaned by a bluestreak cleaner wrasse (Labroides dimidiatus) at Monad Shoal off Malapascua Island, Philippines
Common buzzard (Buteo buteo) landing on prey, wings open, Scotland, UK, September 2016
Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. August
Mountain hare (Lepus timidus) in summer pelage amongst heather, Cairngorms National Park, Scotland, UK, August