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Panamic cushion sea star (Pentaceraster cumingi) detail. Huatulco Bays National Park, Oaxaca state, Mexico, Pacific Ocean. November
Broadleaf woodsorrel (Oxalis latifolia) in flower, close up. Milpa Alta forest, outskirts of Mexico City, Mexico. August
Golden-edged owl-butterfly (Caligo uranus), butterfly captured for the collectors market, wing detail. Montes Azules Biosphere Reserve, Chiapas state, Mexico
Harlequin beetle (Acrocinus longimanus) headshot close up. Los Tuxtlas Biosphere Reserve, Mexico
Common clownfish (Amphiprion ocellaris) in the tentacles of its host, Magnificent sea anemone (Heteractis magnifica), on a coral reef, Bitung, North Sulawesi, Indonesia, Molucca Sea
Velvet swimming crab (Necora puber) male, in defensive posture, with wreck of the Rosalie in background, Weybourne, Norfolk, England, UK, North Sea
Nudibranch (Chromodoris annae) on a coral reef, Bitung, North Sulawesi, Indonesia, Molucca Sea
Orange icing sponge (Mycale laevis) growing on Mountainous star coral (Orbicella faveolata) at an early stage of growth, Cayman Islands, Caribbean Sea
Pair of Nudibranchs (Flabellina exoptata) just prior to mating, the mating apparatus of the one on left is visible, Bitung, North Sulawesi, Indonesia, Molucca Sea
Nudibranch (Phyllidia varicosa) portrait on white background, Bitung, North Sulawesi, Indonesia
Yellowmargin moray (Gymnothorax flavimarginatus) peering out from Digitate leather coral (Sinularia sp)Yellowmargin moray (Gymnothorax flavimarginatus) peering out from Digitate leather coral (Sinularia sp.) on a reef, Ari Atoll, Maldives, Indian Ocean
Hawksbill turtle (Eretmochelys imbricata) feeding on a coral reef, Ari Atoll, Maldives, Indian Ocean. Critically endangered
Roundhead parrotfish (Chlorurus strongycephalus) feeding on coral and algae growing on a reef, South Male Atoll, Maldives, Indian Ocean
Bicolor parrotfish (Cetoscarus bicolor) male, scales detail, Sharm El Sheikh, Sinai, Egypt, Red Sea
Two Tawny nurse sharks (Nebrius ferrugineus) on seabed at dusk, Vavuu Atoll, Maldives, Indian Ocean
Warty frogfish (Antennarius maculatus) yawning in its hiding place amongst white sponges, Bitung, North Sulawesi, Indonesia, Molucca Sea
Asian elephant (Elephas maximus indicus) male, eye detail through foliage, Bardia National Park, Terai, Nepal
Kelp greenling (Hexagrammos decagrammus), female, resting on bed of kelp (Laminariales), Prince William Sound, Alaska, USA, Pacific Ocean
Lion's mane jellyfish (Cyanea capillata) with commensal Crested sculpin (Blepsias bilobus) sheltering amongst stinging tentacles, Prince William Sound, Alaska, USA, Pacific Ocean
Galapagos shark (Carcharhinus galapagensis), listed as potentially dangerous, at sea surface, with nictitating membrane covering its eye, Hawaii, Pacific Ocean
Lemon shark (Negaprion brevirostris) swimming with Remoras (Echeneidae), West End, Grand Bahamas, Atlantic Ocean
Red tailed hawk (Buteo jamaicensis) juvenile female, head portrait, Texas, USA. April
Ants (Formicidae) crawling along cactus spines to escape floodwater, Texas, USA. June
Prickly pear cactus (Opuntia sp. ) flower covered in water droplets, close up, Texas, USA. AprilPrickly pear cactus (Opuntia sp.) flower covered in water droplets, close up, Texas, USA. April
Western diamondback rattlesnake (Crotalus atrox) young, coiled up on desert floor at dusk, flicking tongue, Texas, USA. March
Dung beetle (Scarabaeidae) rolling a ball of dung, Texas, USA. May
Katydid (Tettigoniidae) on Prickly pear (Opuntia sp. ) flower, Texas, USA. MayKatydid (Tettigoniidae) on Prickly pear (Opuntia sp.) flower, Texas, USA. May
Alcedo giant tortoise (Chelonoidis vandenburghi) resting during dry season, Alcedo Volcano, Isabela Island, Galapagos, Ecuador
Detail of hind toes of Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
Close up of Galapagos hawk (Buteo galapagoensis) in immature plumage, flying, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
Scarce swallowtail butterfly (Iphiclides podalirius) landing on a flower, nr Orvieto, Italy. June
Pair of European bee-eaters (Merops apiaster) with courtship offering of insect prey, Pusztaszer, Kiskunsagi National Park, Hungary
Arctic hare (Lepus arcticus groenlandicus) portrait, Ittoqqortoomiit, Northeast Greenland. February
Costa's hummingbird (Calypte costae) male, portrait, showing purple iridescence of head and neck feathers, Coachella Valley, California, USA
Painted frogfish (Antennarius pictus) portrait, Sipidan, Malaysia, Celebes Sea
Close up of Konik wild horse (Equus ferus caballus) snout, Meinerswijk nature reserve, near Arnhem, the Netherlands. May
Portrait of Konik wild horse (Equus ferus caballus), Meinerswijk nature reserve, near Arnhem, the Netherlands. June
Two Konik wild horses (Equus ferus caballus) foals nuzzling, Meinerswijk nature reserve, near Arnhem, the Netherlands. June
Male Highland Great diving beetle (Dytiscus lapponicus) on surface of water, Darnaway Estate, Highlands, Scotland, UK. May. Focus stacked. Captive
Lady's slipper orchid (Cypripedium calceolus) in flower, Swiss Alps, Switzerland. June
African wild dog (Lycaon pictus) alpha female, interacting with two of her pups, aged 4 weeks, Okavango Delta, Northern Botswana, Africa
Greater spearwort (Ranunculus lingua) flower and developing seedhead in the margins of Kenfig Pool, Kenfig National Nature Reserve, Glamorgan, Wales, UK, July
Striped hawkmoth (Hyles livornica) resting on a flower, caught using a MV light trap, Umbria, Italy. July
Red cabbage bug (Eurydema ornata) with a Shield bug (Nezara viridula) side by side on a leaf, Umbria, Italy. September
Crab spider (Synaema globosum) on a Poppy (Papaver sp. ) flower, Sibillini, Umbria, Italy. MayCrab spider (Synaema globosum) on a Poppy (Papaver sp.) flower, Sibillini, Umbria, Italy. May
Broad-bordered bee hawkmoth (Hemaris fuciformis) nectaring on Lavender (Lavandula sp. ) flower, Orvieto, Umbria, ItalyBroad-bordered bee hawkmoth (Hemaris fuciformis) nectaring on Lavender (Lavandula sp.) flower, Orvieto, Umbria, Italy. July
Fly orchid (Ophrys insectifera) in flower, Sibillini, Umbria, Italy. May
Monkey orchid (Orchis simia) in flower, close up, Torre Alfina, Lazio, Italy. May