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Cup fungus (Cookeina sp) Danum Valley, Sabah, Borneo
Yew (Taxus baccata) berries. Strumpshaw Fen, Norfolk, UK, September
Black Grouse (Tetrao tetrix) displaying with breath vapor at dawn, Utajarvi, Finland, April
Water lily (Nymphaea rudgeana) leaves viewed from below in a black water creek. Suriname, October
Hydromedusa / Trachymedusa (Crossota sp) from approx 2700m, mid Atlantic ridge, June 2010
RF- Hyacinth macaw (Anodorhynchus hyacinthinus) in its nest hole. Pousada Aguape Lodge, Mato Grosso do Sul State, Brazil. September
Cheetah (Acinonyx jubatus) cub aged 6 months yawning, Masai-Mara Game Reserve, Kenya. Vulnerable species
Konik Wild Horse (Equus ferus caballus) yawning. The Netherlands, November
Boer domestic goat (Capra hircus) waiting to be fed, Norfolk, UK, September
Artic fox (Vulpes lagopus) portrait, captive, Norway, February
Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards
Deer mouse (Peromyscus maniculatus) on giant puffball mushroom, watching mosquito in the moonlight. Blackfoot Valley, Western Montana, USA, July
Cloud iridescence forming above a cumulonimbus cloud, caused by light refraction through individual water droplets and ice crystals at high altitude, Sierra Leone
Lemur leaf frog (Hylomantis lemur) mating pair, captive, Critically endangered, from Central America. Please credit Michael Kern / Zoo Atlanta
Common frog (Rana temporaria) sheltering from rain under toadstool (Macrolepiota procera) the Netherlands
Herring Gull (Larus argentatus) close up of juvenile in flight, Flatanger, Norway, October
RF - Siberian tiger (Panthera tigris altaica) in snow, captive
Split level photo of an American crocodile (Crocodylus acutus) floating at the surface over a shallow seagrass meadow, close to mangroves
Sunflower (Helianthus annuus) Spain
Scots pine (Pinus sylvestris) mature tree in evening light, Abernethy Forest RSPB Reserve, Cairngorms National Park, August
Mustang / wild horses, second year cremello colt Claro play fighting with bay, McCullough Peak herd, Wyoming, USA, July 2009
Grey angelfish (Pomacanthus arcuatus) portrait, Georgetown, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Rock beauty angelfish (Holacanthus tricolor) pair at dusk performing their courtship dance with the larger male circling the female, Georgetown, Grand Cayman, Cayman Islands, British West Indies
Schoolmaster snapper (Lutjanus apodus) above a coral reef, Georgetown, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February
Briard / Berger de Brie, head portrait with plaited hair between eyes
Gloucester old spot piglets (Sus scrofa domestica) suckling at the Cotswold Farm Show, Cirencester, UK, July 2008
Rothschilds giraffe (Giraffa camelopardalis rothschildi) close up of young calf skin pattern, captive
Grevy's zebra (Equus grevyi) close up, captive
Red fox (Vulpes vulpes) close-up of half of face, captive
Black bear (ursus americanus) cub standing on hind legs, Yellowstone National Park, USA, June
Blue Shark (Prionace glauca) near sea surface and divers in background. Santa Maria, Azores, September
Demoiselle crane (Anthropoides virgo) portrait, captive
Icebergs off the Antarctic Peninsula, Antarctica, February 2009, Taken on location for BBC Frozen Planet series
Aphelandra (Acanthus sp) flowering. Oahu, Hawaii, February
Common buckeye butterfly (Junonia coenia) feeding on Ironweed (Vernonia altissima) flower, North Guilford, Connecticut, USA, August
Ocellated eagle ray (Aetobatus ocellatus), underside, Natal Coast, South Africa
Black blotched pufferfish (Diodon liturosus) inflated with water at night as defensive behaviour, Sipadan Island, Borneo, Malaysia
Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October
Leaf cutter ant (Atta sp) female worker carrying leaf to nest, with a smaller minor ant riding. The minors job is to protect her sister from parasitic flies. Costa Rica, December
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
Hoopoe (Upupa epops) in flight, Hungary. Multi flash, long exposure. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition
Green crowned brilliant hummingbird (Heliodoxa jacula) male taking off to feed at feeder in garden, another waiting perched on lichen covered branch, Costa Rica, January
Red Fox (Vulpes vulpes) vixen on a misty day in woodland, Black Forest, Germany, July. Runner-up in Animal Portraits category, Wildlife Photographer of the Year competition 2011
Bigeye Scad (Selarcrumenophthalmus). Misool, Raja Ampat, West Papua, Indonesia
Pipevine Swallowtail Butterfly (Battus philenor) pair mating. Sinton, Corpus Christi, Coastal Bend, Texas, USA, June
Red toy Poodle dog, and Ragdoll-cross kitten, 5 weeks
Tabby kitten, 5 months, standing up with raised paws
