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Grass snake (Natrix natrix) flicking tongue, Hampstead Heath, London, England, UK, August
Mountain hare (Lepus timidus) on snow, Cairngorms National Park, Scotland. January
Red Squirrel (Sciurus vulgaris) adult sitting and feeding in snowfall. Cairngorms National Park, Scotland, UK, March
RF- Common frog (Rana temporaria) portrait, Cornwall, UK, January
Water vole (Arvicola amphibius) standing on hind legs sniffing blackberry, Kent, UK, September
Eyes of Margined sole fish (Brachirus heterolapis) on its camouflaged body. Saonek Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean. Dampier Strait
Reticulate stargazer (Uranoscopus sp. ) emerging from the sandReticulate stargazer (Uranoscopus sp.) emerging from the sand. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Large female Southern stingray (Hypanus americanus) in shallow water, under storm clouds, Stingray City, Grand Cayman. Stingray City is feeding site for stingrays and bait was used on this shoot
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Butterfish (Pholis gunnellus) portrait. Gardur, south west Iceland. North Atlantic Ocean
Red Irish lord (Hemilepidotus hemilepidotus) hides on the sebed, with Quillback rockfish (Sebastes maliger) and copper rockfish (Sebastes caurinus) behind in the bull kelp forest
Blue sharks (Prionace glauca) cruise beneath the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Close up of pattern of the cerata of a nudibranch (Janolus savinkini). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Coleman shrimp (Periclimenes colemani) close up, on a fire urchin (Asthenosoma varium). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
RF- Peregrine Falcon (Falco peregrinus) landing on moorland with its wings spread. Scotland, UK, March
Purple short tentacle mushroom coral (Fungia sp. ) close up of polyps. Lembeh StraitPurple short tentacle mushroom coral (Fungia sp.) close up of polyps. Lembeh Strait, North Sulawesi, Indonesia
Foxglove (Digitalis purpurea) flower detail, Devon, UK. June
Crocus flower (Crocus sp) stigma - close up
RF - Early Purple Orchid (Orchis mascula) in flower, Broxwater, Cornwall, UK. May 2016
Pasque flowers (Pulsatilla vulgaris) in rain, Lorraine, France, April. 3rd Prize in the Other Nature Topics category of Melvita Nature Images Awards 2013
Lavender (Lavandula angustifolia) fields, Valensole Plateau, Alpes Haute Provence, France, June
RF- Little blue heron (Egretta caerulea) and Great egret (Ardea alba) in lagoon at sunrise. Fort Myers beach, Gulf Coast, Florida, USA, March
Stone canyon jasper formed with cryptocrystalline quartz, California, USA
Hedge Parsley seed head (Torilis japonica) silhouetted at sunset, Norfolk, UK, November
Gorgonian / Fan coral (Solenocaulon akalyx), close up, West Papua, Indonesia
Close-up of the polyps out of a coral tree (Tubastraea coccinea) at night, Baja California peninsula, Mexico. Sea of Cortez
RF- Chanterelle mushroom (Cantharellus cibarius), gill detail
Orange cup coral (Tubastrea coccinea) Malpelo Island National Park, UNESCO World Heritage Site, Colombia, East Pacific Ocean
Red Admiral Butterfly (Vanessa atalanta) on Michaelmas daisy flowers. Dorset, UK, October 2012
Coral clam (Pedum spondyloideum) close up, Maldives, Indian Ocean
Two Great egrets (Ardea alba) wading, silhouetted at dawn, Keoladeo National Park, Bharatpur, Rajasthan, India
Great Tit (Parus major) perched on branch with yellow from autumn trees in the background. Southern Norway, October
Endive Pellia liverwort (Pellia endiviifolia) in centre growing through Common Liverwort (Marchantia polymorpha), the latter bearing cups containing gemmae (used in asexual reproduction)
Sword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devil sSword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devils Trumpet Flower (Datura sp.). Yanacocha montane cloud forest, Ecuador
RF- Pool Frog (Rana lessonae) sitting on white lily pad, Danube delta rewilding area, Romania
RF- Boulder star coral (Montastrea annularis) showing fluorescent green coloration, photographed under blue light at night, to excite fluorescent pigments in the coral, East End, Grand Cayman
Eastern green mamba {Dendroaspis angusticeps} skin detail, captive, from East Africa
Peacock (Pavo cristatus feather close up showing iridescence at 10x magnification
Close-up of Hyacinth Macaw (Anodorhynchus hyacinthinus feathers, Brazil
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
RF - Long-spined sea urchin (Astropyga radiata). Lembeh Strait, North Sulawesi, Indonesia
Gorgonian coral (Acanthogorgia sp. ) Similan Islands, Andaman Sea, ThailandGorgonian coral (Acanthogorgia sp.) Similan Islands, Andaman Sea, Thailand
Peacock butterfly (Inachis io) feeding on Goat Willow catkins (Salix caprea), an important food source for pollinators in early spring. Peak District National Park, UK. April
RF- Close up of Leaves of Red Oak (Quercus rubra) in autumn. October
Big-legged Bug or Squash Bug (Coreidae) on Heliconia Flower. Yasuni National Park, Amazon Rainforest, Ecuador. South America
Cup fungus (Cookeina sp) Danum Valley, Sabah, Borneo