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Green turtle (Chelonia mydas) from underneath swimming through the surface, Mayotte Island, Comores, Indian Ocean
Wild gladiolus (Gladiolus illyricus) flowers, Menorca, Balearic Islands, Spain, Europe
RF - Granular poison frog (Oophaga granulifera). Osa Peninsula, Costa Rica. Vulnerable species
Little blue / fairy penguin (Eudyptula minor) walking on pathway to nesting burrows, Neck Game Reserve, Bruny Island, Tasmania, Australia, December
Gentoo Penguin (Pygoscelis papua) jumping out of the sea, Cuverville Island, Antarctic Peninsula, Antarctica
King Penguin (Aptenodytes patagonicus) adult standing amongst colony of chicks, Gold Harbor, South Georgia, South Atlantic Islands
Pacific hatchetfish (Argyropelecus affinis) portrait
Deepsea squid from midwater catch 195-498m, Mid-Atlantic Ridge, North Atlantic Ocean
Deepsea squid (Histioteuthis sp) from between 188m / 617ft and 507m / 1, 663ft depth, nightDeepsea squid (Histioteuthis sp) from between 188m/617ft and 507m/1, 663ft depth, night, Mid-Atlantic Ridge, North Atlantic Ocean
Shelled Pteropod / Sea butterfly {Diacria trispinosa} with egg string attached, from between 188 and 507m, Mid-Atlantic Ridge, North Atlantic Ocean
Deepsea Medusa jellyfish (Colobonema sp), Mid-Atlantic Ridge, North Atlantic Ocean
RF - Harts tongue fern (Phyllitis scolopendrium) leaf unfurling and creating the shape of a heart, Cornwall, UK, May
RF- Gannets (Morus) displaying, Great Saltee, County Wexford, Ireland. May
Koala (Phascolarctos cinereus) mother with joey aged eight months, Queensland, Australia, captive
Gannet (Morus bassanus) breeding pair during part of their elaborate courtship ritual. Saltee Islands, Republic of Ireland, May
Giraffe (Giraffa camelopardalis) female bending down to calf, Okavango Delta, Botswana
Unravelling Fern fronds (Unknown species). Near Nepenthes Field Camp, mid-altitude montane forest. Centre of Maliau Basin, Sabahs Lost World, Borneo
Feral domestic rabbit (Oryctolagus cuniculus) juvenile running with dead leaf in mouth, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Booby (Sula nebouxii) looking at camera underwater, Galapagos Island
Red-legged Seriema (Cariama cristata) portrait, captive, Apenheul Park, Netherlands
Spectacled caiman (Caiman crocodilus) Rio BaiAia Bonita, Bonito, Mato Grosso do SulSpectacled caiman (Caiman crocodilus) Rio Baiia Bonita, Bonito, Mato Grosso do Sul, Brazil
Birch Sawfly (Cimbex femoratus) feeding on a leaf. August, UK
Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills, Lake Kerkini, Greece, February. Highly Commended, 2011 Wildlife Photographer of the Year competition
Four Mallard (Anas platyrhynchos) dabbling for food in icy water. The Netherlands, January
Basset hound, portrait, resting head on back of car, eyes half closed, UK
Fulmar (Fulmarus glacialis) rear view of bird hanging in air over steep cliffs, Shetland Islands, Scotland, UK. HIGHLY COMMENDED - Animals in their environment - Wildlife Photographer of the Year
Head portrait of Highland cow, Scotland, with tiny frond of bracken at corner of mouth, UK
Spring snowflake flower (Leucojum vernum) with bokeh effect, Vosges, France, March
RF - Blue waterlily (Nymphaea caerulea) close up into centre of flower. Occurs in Africa and Asia. Botanic Garden Meise, Belgium, August
RF - Close up of Greater stitchwort (Stellaria holostea) flower, Cornwall, UK. April
Common Bird s-foot-trefoil (Lotus corniculatus) Nordtirol, Austrian Alps. June
Giant Water Lily (Victoria amazonica) flower in a lagoon at Porto Jofre, off the Cuiaba River, northern Pantanal, Mato Grosso State, Brazil, South America
Graylag gosling (Anser anser) with parent, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. April
Japanese macaque (Macaca fuscata fuscata) mother with rare white furred baby, Jigokudani Valley, Nagano Prefecture, Japan. June
RF- European hedgehog (Erinaceus europaeus) orphan, Jarfalla, Sweden. August
Baby Olive Ridley sea turtle (Lepidochelys olivacea) struggles against the swell to swim away from the beach where it hatched, Ostional, Costa Rica. October
Close-up of an eye of a Sumatran tiger (Panthera tigris sumatrae), captive, occurs in Sumatra, Indonesia
RF- Juvenile Sumatran tiger (Panthera tigris sumatrae), aged four months, captive, occurs in Sumatra, Indonesia
Bengal Tiger (Panthera tigris tigris) captive, with leaves digitally added
Bengal Tiger (Panthera tigris) sub-adult, approximately 17-19 months old, amongst forest vegetation. Endangered. Bandhavgarh National Park, India. Non-ex
Bengal Tiger (Panthera tigris tigris) young cub, Pench National Park, Madhya Pradesh, India, taken on location for Tiger - Spy in the Jungle July 2008
RF- Siberian tiger (Panthera tigris altaica) young cub, lying down, looking away, captive
RF- Male Bengal Tiger (Panthera tigris tigris) face portrait in undergrowth, Kanha National Park, Madhya Pradesh, India. Endangered species
RF- Bottlenose dolphin (Tursiops truncatus) swimming in a spiral movement, Sandy Ridge, Little Bahama Bank. Bahamas
Blackbird (Turdus merula) and falling snow. Leicestershire, UK, February
Rock Martin (ptyonoprogne fuligula)in flight, Israel
Detail of White Witch Moth (Thysania agrippina) in cloud forest, Manu Biosphere Reserve, Amazonia, Peru
Palm (Areaceae) leaves silhouetted in mist at dawn, Royal Chitwan National Park, Nepal
