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Blue wildebeest (Connochaetes taurinus), migrating herd silhouetted on the horizon at sunset, Masai Mara Game Reserve, Kenya
Jocotoco antpitta (Grallaria ridgelyi) flagship species for Jocotoco Conservation Foundation and Tapichalaca Biological Reserve, Province Zamora-Chinchipe, Ecuador
Yellowfronted canaries (Crithagra mozambica) at waterhole, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa. June
Leatherback turtle (Dermochelys coriacea) female returning to sea after laying eggs, Trinidad and Tobago, April
Spectacled weaver (Ploceus ocularis) bathing, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa, June
North American Bison (Bison bison) coated in frost standing on snow, Yellowstone National Park, Wyoming, USA, January
Globe Thistle (Echinops ritro), dead in winter landscape with Alpine ibex (Capra ibex) silhouet behind, Gran Paradiso national park, Italian Alps, Italy, January
White-throated kingfisher (Halcyon smyrnensis) with Black drongo (Dicrurus macrocercus) Ranthambore National Park, Rajasthan, India
Great tit (Parus major) sitting on Hazel (Corylus) with catkins, Loire Atlantique, France, January
Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean
Cley Beach with scattering of light snow, and boats, Norfolk, England, UK, December 2010
Dalmatian pelicans (Pelecanus crispus) squabbling over fish, Lake Kerkini, Greece, February
Dahlias in flower, seen through foggy cottage window, Ringshall, Hertfordshire, England, UK, September
Ruff (Philomachus pugnax) males fighting at lek, Varanger, Norway, May 2016
Cloud forest scene in the Monteverde Cloud Forest Reserve, Costa Rica, February 2015
Eastern chipmunk, (Tamias striatus) in New Brunswick, Canada. June 2014
Coquerels sifaka (Propithecus coquereli) curious individual up in tree, Ankarafantsika National Park, Madagascar
White-breasted Fruit-Dove (Ptilinopus rivoli) male at fruiting Shefflera tree. Huon Peninsula, Morobe Province, Papua New Guinea
Rainforest and valley in the Sarawaget range, Huon Peninsula, Papua New Guinea. November 2006
Katydid (Tettigoniidae) with Giraffe (Giraffa camelopardalis) in the background, Masai Mara National Park, Kenya
Broken window overgrown with ferns, Kells Seaside Area, Ring of Kerry, Iveragh Peninsula, County Kerry, Ireland, Europe. September 2015
Moonlight over the Skellig Islands, Puffin Island, County Kerry, Ireland, Europe. September 2015
Sea chamomile (Anthemis maritima), Sea stock (Mattiola sinuata), and Shrubby Everlasting (Helichrysum stoechas) flowers in sand dunes, Camargue, France. May
Stellers sea eagle (Haliaeetus pelagicus) in flight with mountains behind, Hokkaido Japan February
Aurora Borealis and coastline at Hornstrandir, Iceland. April 2016
Water vole (Arvicola amphibius) feeding on aquatic plant, Kent, UK May
Pine marten (Martes martes) running across branch, Black Isle, Scotland, UK. October, taken by remote camera trap
Bird nesting cliffs at Haeavikurbjarg, Hornvik, Hornstrandir, Iceland. July
Tree covered in ice and frost during snowfall with heavy wind, Hohneck mountain, Vosges, France, January
Great grey owl (Strix nebulosa) head portrait, Kuusamo, Finland, March
Cormorant (Phalacrocorax carbo) swallowing caught fish, Hungary January
Japanese / Red-crowned crane (Grus japonicus) three coming into land, Hokkaido Japan February
Black stork (Ciconia nigra) beak submerged fishing, Hungary May
River stingray (Potamotrygon sp) Formoso River, Bonito, Mato Grosso do Sul, Brazil
Common reed bunting (Emberiza schoeniclus) attacking Common cuckoo (Cuculus canorus), female, Braunschweig, Lower Saxony, Germany, May
Whooper swans (Cygnus cygnus) fighting, Finland, April
Whooper swan (Cygnus cygnus) pair in flight in snowy landscape, Finland, November
Grey seal (Halichoerus grypus) young male plays and gapes, showing his teeth. Lundy IslandGrey seal (Halichoerus grypus) young male plays and gapes, showing his teeth.Lundy Island, Devon, UK, Bristol Channel, August
Perito Moreno Glacier, Los Glaciares National Park, Santa Cruz, Patagonia, Argentina. February 2010
Thonga Beach on the coast of Indian Ocean, Maputuland, KwaZulu-Natal, South Africa, January
Crocodilefish (Cymbacephalus beauforti) close up detail of the retractable eye-lashes, Mabul Island, Sabah, Borneo, Malaysia Celebes Sea
Soft corals (Dendronephthya hemprichi) growing in very shallow water in the shade provided by a jetty, while a school of Silversides (Atherinomorus lacunosus) circle. Berenice Jetty, Aqaba, Jordan
Ice calving from Perito Moreno Glacier, Los Glaciares National Park, Santa Cruz, Patagonia, Argentina. February 2010
Mountainous landscape in Torres del Paine National Park, with mountains behind, Chile, June 2015
Silversides (Atherinomorus lacunosus) school just below surface in evening light, Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
Polar bear (Ursus maritimus) resting on sand, Kaktovik, Alaska, USA, September
Close up of Red deer stag (Cervus elaphus) with antlers covered in velvet in spring, Daun, Vulkaneifel district, Rhineland-Palatinate, Germany, June. Captive
Cape buffalo (Syncerus caffer) drinking at waterhole at night, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa, September