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Lion (Panthera leo) side profile of head, at night, Central Kalahari Game Reserve, Botswana, digitally enhanced
Head portrait of grey almost white Andalusian mare running in snow, Berthoud, Colorado, USA. January
Asiatic elephants (Elephas maximus), dust bathing at dawn. Jim Corbett National Park, India
Kottigehara dancing frog (Micrixalus kottigeharensis), dancing frog name given because of foot flagging behavior between competing males. Endemic to Western Ghats. Agumbe, Karnataka, India
Jayarami Bush frog (Raorchestes jayarami), inflating vocal sac, calling to attract mate, Endemic to Western Ghats. Anamalai Wildlife Sanctuary, India
Ocelot (Leopardus pardalis) portrait. camera trap image, Nicoya Peninsula, Costa Rica
Polar bear (Ursus maritimus) aerial view of sow with cub along the Arctic coast in summer, near Point Barrow, Alaska, USA. July
Black-crested titmouse (Baeolophus bicolor), adult perched on icy branch of Possum Haw Holly (Ilex decidua) with berries, Hill Country, Texas, USA. February
Sea stack and Talisker Beach, Isle of Skye, Inner Hebrides, Scotland, UK, September 2013
Scots pine (Pinus sylvestris) forest in evening light, Abernethy, Cairngorms National Park, Scotland, UK, June
Bison (Bison bison) close up head portrait with snow on fur, Yellowstone National Park, USA, February
Pine marten (Martes martes) leaping, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Animal Behaviour category of the BWPA (British Wildlife Photography Awards) Competition 2016
Gannet (Sula bassana) in water, Shetland, UK. July
Kestrel (Falco tinnunculus) male, Hampstead Heath, England, UK
Red kite (Milvus milvus) in a dive, Gigrin Farm, Rhayader, Wales, UK April
Studio image of multiple Nudibranch species. Gulen, Norway. North East Atlantic Ocean. Digital composite
School of Blacksmith fish (Chromis punctipinnis) swim through a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
Napoleon wrasse (Cheilinus undulatus) portrait. Red Sea, Egypt. October
Cirrus clouds displaying wind shear, Brechin, Scotland, UK, September
Monte Perdido massif and Las Tres Sorores (The Three Sisters) Ordesa National Park, Aragon, Huesca, Spain
Portrait of an African elephant (Loxodonta africana) on Selinda Reserve, Botswana
Common dolphin (Delphinus delphis) reflection as its swimming on the surface of the ocean, Eastern Pacific, March
Sperm whales (Physeter macrocephalus) large group gathering together, rubbing together to shed skin, interacting and defecating. Indian Ocean, March
Willow grouse (Lagopus lagopus) camouflaged against snow, Inari, Kiilopaa, Finland, February
Bay of Laig, Isle of Eigg towards Isle of Rum, Inner Hebrides, Scotland, UK, April 2014
Sedge warbler (Acrocephalus schoenobaenus) singing with bokeh effect in background, Norfolk, England, UK, May. Winner of the Birds in Environment Category of the Bird Photographer of the Year 2015
Natural ice sculpture floating at sea in Svalbard, Norway
Atlantic puffin (Fratercula arctica) swimming on the ocean surface, Svalbard, Norway, June
Panoramic landscape of Cerredo mountain at sunset, Castro Urdiales, Cantabria, Spain
Eurasian / European badger (Meles meles) outside urban sett behind wall with graffiti, Bristol, UK May
Common / Northern minke whale (Balaenoptera acutorostrata) surfacing, Pacific coast of Baja California, Mexico, February
Grayling (Thymallus) under Ice, Lake Baikal, Siberia, Russia
Royal spoonbill (Platalea regia) preening and shaking head feathers, Victoria, Australia
Grey heron (Ardea cinerea) in rain with another heron behind, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Two Buzzards (Buteo buteo) fighting, Lake Csaj, Kiskunsagi National Park, Hungary. February
Cormorants (Phalacrocorax carbo) squabbling over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Two Burrowing owls (Athene cunicularia) fledgling with adult about to take off, Pantanal, Brazil
Grey herons (Ardea cinerea) fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. February
Shadowfin soldierfish (Myripristis adusta) West Papua, Indonesia
Shrimpfish / Razorfish (Aeoliscus punctatus / strigatus) have adapted to a vertical posture and swim with their head downward. Lembeh, North Sulawesi, Indonesia
Wide angle view of Demoiselle cranes (Anthropoides virgo) landing in chugga ghar (bird feeding enclosure), Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) flock taking flight during winter migration. Khichan, Western Rajasthan, India. February 2015
Red-and-green macaws (Ara chloropterus) perched and in flight over Buraco das Araras, the Sinkhole of the Macaws, Jardim, Mato Grosso do Sul, Brazil. September
Early morning shafts of light through deciduous forest. Bandhavgarh National Park, India
Sun beam at sun rise over the Grand Canyon of the Yellowstone, Yelllowstone National Park, Wyoming, USA. January 2016
Cluster of Burnt or Burnt-tip Orchids (Neotinea ustulata) in a Alpine meadow. Tyrol, Austria