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Great egrets (Ardea alba) sky pointing, sizing each other up before fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Plains zebra (Equus quagga) close up of face, Kariega Game Reserve. South Africa
Flock of Mallard ducks (Anas platyrhynchos) taking off, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. December
Mud-covered face of bull African buffalo (Syncerus caffer) with a cateract in one eye, on the banks of the Chobe River, Chobe National Park, Botswana
Western diamond-backed rattlesnake (Crotalus atrox) Texas, USA, April
Great egret (Ardea alba) and Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
European robin (Erithacus rubecula) low angle view near church, Bristol, UK. January
Mountain landscape reflected in lake, Basongcuo National Park, Qinghai-Tibet Plateau, Tibet, November 2011
Red mason bee (Osmia rufa) female emerging from her nest hole in a drilled log within an insect hotel after provisioning a brood cell with pollen, Gloucestershire garden, UK, April
Common eider (Somateria mollissima), female on her nest with sea behind, Finland, May
Group of Great egrets (Ardea alba), Grey heron (Ardea cinerea) and Eurasian spoonbill (Platalea leucorodia) at night with stars and moon. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. May
Common raven (Corvus corax) two adults in flight, Hornoya, Norway, April
Hawfinch (Coccothraustes coccothraustes), male perched on twig, Uto, Finland, May
Milky Way rising over the south wall of the Grand Canyon, Arizona, USA. November 1 2015. Composite image
European Chubb (Squalius / Leuciscus cephalus) in cave, Plitvice Lakes National Park, Croatia. November
Scorpion (Heterometrus sp) fluorescing in ultraviolet light at night on forest floor, Danum Valley, Sabah, Borneo
Andromeda (M31) with its two syster galaxies (M32 and M110) Eastern Colorado. 8 November 2015. Taken with digital focus stacking
Underwater view of the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
Bealach na Ba Pass, Wester Ross, Scotland, UK, November 2015
European hamster (Cricetus cricetus) male, in underground burrow, captive
Namib sand snake (Psammophis namibensis) in sand dunes, Swakopmund, Erongo Region, Namibia
Orange anemonefish (Amphiprion sandaracinos) in its host Mertens carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea
Flame anthias (Pseudanthias ignitus) displaying to the more drab female (above) during courtship in the evening. Similan Islands, Thailand. Andaman Sea, Indian Ocean
Great hammerhead shark (Sphyrna mokarran) female at night with Bar jacks (Caranx ruber), South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Ol Doinyo Lengai, an active volcano held in high esteem and known locally as The Mountain of God by the Masai, with people and huts on the savannah below, Rift Valley, Tanzania
Puma (Puma concolor) in high altitude habitat of Torres del Paine National Park, Chile
Alpine ibex (Capra ibex) adult male in deep snow on a ridge with young during snowfall, Gran Paradiso National Park, the Alps, Italy. January
Head portrait of black Andalusian stallion and mare meeting for the first time in southern Spain, Europe
Masked tree frog (Smilisca phaeota) El Arenal region, Costa Rica
Night-blooming cereus cactus (Acanthocereus tetragonus), flower bud opening at night, Texas, USA. Sequence 7 of 7. August
Blue cheeked bee eater (Merops persicus) two squabbling, Oman, April
Malabar pit viper (Trimeresurus malabaricus), green colour morph. Agumbe, Karnataka, Western Ghats, India. Endemic
King cobra (Ophiophagus hannah), low wide angle perspective Agumbe, Karnataka, Western Ghats, India. Vulnerable species
Siskin (Carduelis spinus) male in flight preparing to land, Scotland, UK, May
Red Deer (Cervus elaphus) herd in forest in snow, Cairngorms National Park, Scotland, UK. December
Polar bear (Ursus maritimus) sow with two cubs walking past a pile of Bowhead whale (Balaena mysticetus) bones, Kaktovik, Barter Island, 1002 Area, Arctic National Wildlife Refuge, North Slope
Polar bear (Ursus maritimus) spring cub sitting upright on hind legs balancing, Bernard Spit, 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Julia heleconia (Dryas iulia) butterfly on head of Yacare caiman (Caiman yacare). Butterflies often land on caimans head to drink the salt from its eyes. Pantanal, Brazil
Aerial view of two Gemsbok (Oryx gazella) in sand dunes, Namib Desert, Namibia
Spotted catshark (Scyliorhinus canicula) just hatched from egg case that is on Seafan (Paramuricea clavata), Ullastres seamount, Costa Brava, Llafranch, Catalunya, Spain
Blue shark (Prionace glauca) seen from above with pilot fish, Azores Islands, Portugal, Atlantic Ocean
Giant moray eel (Gymnothorax javanicus) opening its mouth to breathe, at night on the barge wreck, Gubal Island, northern Red Sea
Spectacled caiman (Caiman crocodilus) in water at night, Mato Grosso, Pantanal, Brazil
Pink-footed goose (Anser brachyrhynchus) flock flying in mist, Cromarty Firth, Highlands, Scotland, UK April
Curlew (Numenius arquata) roosting on saltmarsh, Cromarty Firth, Scotland, UK April
Septarian concretion or (septarian nodules) from Morocco. Dense mass formed by precipitation of mineral cement in sedimentary rock or soil which is later cracked and filled with other material
Orbicular jasper, ocean jasper from Madagascar, a variety of jasper which contains spherical inclusions