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Cape buffalo (Syncerus caffer) drinking at waterhole at night, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa, September
Emergence of Mexican free-tailed bats (Tadarida brasiliensis) from Bracken Cave, Texas. Bracken Cave is the summertime home of over 15 million bats
European beewolf (Philanthus triangulum) flying, carrying paralysed bee to nest, Budapest, Hungary
Alpine ibex (Capra ibex) close-up crop head portrait of an adult male, Gran Paradiso National Park, the Alps, Italy. January
Gannet (Morus bassanus) nearest out of focus, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Red squirrel (Sciurus vulgaris) half way up Scots pine (Pinus sylvestris) Black Isle, Scotland, UK. October
Split level view of Blue shark (Prionace glauca) at surface at sunset, Azores Islands, Portugal, Atlantic Ocean
Meadow brown (Maniola jurtina) resting on grass in Cohens Field, Hampstead Heath, London, England, UK, June
Ice covered Reindeer (Rangifer tarandus) during storm. Yar-Sale district, Yamal, Northwest Siberia, Russia. April
Snowy owl (Bubo scandiaca) female face close up, Canada February
Cormorant (Phalacrocorax carbo) swallowing perch prey, Hungary, January
Golden-brown mouse lemur (Microcebus ravelobensis), Ankarafantsika National Park, Madagascar
Cley Beach covered in snow, Norfolk. England, UK, January
Ruff (Philomachus pugnax) male displaying, Varanger, Norway, May 2016
Storm clouds over Hornvik, Hornstrandir, Iceland. July 2015
Birch (Betula pendula) and Scots pine (Pinus sylvestris) forest around Loch Affric in dramatic light, Glen Affric, Highlands, Scotland, UK February
Storm waves breaking against cliffs at Svortuloft, Iceland April 2016
Lichens on rock, Torridon, Scotland, UK, November
Waves at Malariff, Iceland April 2016
Painted lichen moth (Hypoprepia fucosa) on autumn leaf, Tuscaloosa County, Alabama, USA September
Implicit arches moth (Lacinipolia implicata) on white background, Tuscaloosa County, Alabama, USA October
Common blue butterflies (Polyommatus icarus) soft focus resting on grass, Vealand Farm Devon Wildlife Trust Reserve, Devon, England, UK. September
Agua Rica leaf frog (Phyllomedusa ecuatoriana) captive, endemic to Agua Rica, Ecuador. Endangered species
Leafcutter ants (Atta sp) colony harvesting pieces of a banana leaf, Costa Rica
Gorgonian sea fans (Melithaea sp. ) with a large school of Pygmy sweepers (ParapriacanthusGorgonian sea fans (Melithaea sp.) with a large school of Pygmy sweepers (Parapriacanthus ransonetti) Andaman Sea, Thailand
Great egret (Ardea alba) group of five at waters edge, with very out of focus egret in foreground. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, February
Little bittern (Ixobrychus minutus) hunting from reed, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, June
Cormorant (Phalacrocorax carbo) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, January
Egytian Goose (Alopochen aegyptiaca) rear view bathing, sings spread, Zimanga Private Game Reserves, South Africa
Pine trees in mist, Ballons des Vosges Regional Natural Park, Vosges, France, October 2014
Greater flamingo (Phoenicopterus roseus) feathers, Pont de Gau Ornithological Park, Camargue, France, March
A misty sunrise over tree silhouette in Deadvlei, Namibia
Raft Spider (Dolomedes fimbriatus) female resting on the surface of a moorland pool, photographed with a fisheye lens to show surrounding habitat. Nordtirol, Austrian Alps, June
Black headed gull (Chroicocephalus ridibundus) taking off, detail of legs and water droplets, Pusztaszer, Hungary, April
Great white egret (Egretta alba) detail of plumage and legs from the front, Pusztaszer, Hungary, April
Beautiful pitviper (Trimeresurus venustus) juvenile, captive occurs in Thailand. Venomous species
King cobra (Ophiophagus hannah) juvenile in threat pose on white background, captive occurs in South Asia. Venomous species
Yellow Labrador retriever puppy, age 9 weeks, sleeping
Red dunes of Sossusvlei, Namibia
Lesser flamingos (Phoeniconaias minor) flock on lake at dawn, Lake Ndutu Tanzania
Chamois (Rupicapra rupicapra) in deep snow, Gran Paradiso National Park, Italy. November. Commended in the Asferico Photography Competition 2016
Standard Poodle in sharp wind on beach dune, Waterford, Connecticut, USA
English Bulldog in daffodils, Waterford, Connecticut, USA
Yellow Labrador retriever romping in fresh snow, Clinton, Connecticut, USA
Red snappers / Two-spot red snappers (Lutjanus bohar) mass shoal of fish spawning, Palau, Philippine Sea. A few black and white snappers (Macolor niger) have crawled into the school to eat the eggs
Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica
Veragua cross-banded treefrogs (Smilisca sordida) mating pair at night, Osa Peninsula, Costa Rica
Thornbug (Umbonia crassicornis) San Jose, Costa Rica
