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Pink anemonefish (Amphiprion perideraion) in Purple magnificent sea anemone (Heteractis magnifica) Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
Steller sealion (Eumetopias jubatus) male charging past the camera. Race Rocks, Victoria, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
Maldive anemonefish (Amphiprion nigripes) shoal swimming in the current flowing over a Magnificent sea anemone (Heteractis magnifica). Baa Atoll, Maldives. Indian Ocean
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon, on coral reef under stormy seas. St Johns Reef. Egypt. Red Sea
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
California sea lions (Zalophus californianus) group playing in the sun in the early morning. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Bigfin reef squid (Sepioteuthis lessoniana) capturing a pelagic shrimp with long red antennae, in midwater, at night. Anilao, Batangas, Luzon, Philippines
Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Common starling (Sturnus vulgaris) flock at dusk engaging in pre-roost display, Dumfries-shire, Scotland, March
Pink anemonefish (Amphiprion perideraion) in a purple Magnificent sea anemone (Heteractis magnifica). Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Ruppells weaver (Ploceus galbula) male building nest, Oman, May
Juvenile Andean mouse opossum (Marmosops impavidus) eating a moth, El Oro, Ecuador
Hourglass treefrog (Dendropsophus ebraccatus) male with inflated vocal sac calling at night, Central Caribbean foothills, Costa Rica
Black skimmer (Rynchops flavirostris) fishing by skimming along water surface, Pantanal, Brazil
Three-striped poison frog (Ameerega trivittata) male carrying tadpoles, Panguana Reserve, Huanuco province, Amazon basin, Peru
Black-striped capuchin (Sapajus libidinosus) using rock as a tool to break open palm nut, Parnaiba Headwaters National Park, Piaui, Brazil. August
Short-beaked echidna (Tachyglossus aculeatus) digging, Tasmania, Australia
Sword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devil sSword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devils Trumpet Flower (Datura sp.). Yanacocha montane cloud forest, Ecuador
Indochinese tiger (Panthera tigris corbetti) looking up, behind screen of leaves, captive, occurs in South East Asia
Male Yellow warbler (Setophaga / Dendroica petechia) in breeding plumage singing amongst oak catkins and leaves, New York, USA, May
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Black Grouse (Tetrao tetrix) displaying with breath vapor at dawn, Utajarvi, Finland, April
Mother and baby Bornean orangutan (Pongo pygmaeus) in trees Tanjung Puting National Park, Borneo-Kalimatan, Indonesia, endangered species
Red deer (Cervus elaphus) stag bellowing during rut, Derbyshire, UK
Musk ox (Ovibos moschatus) portrait of two standing closely, Wrangel Island, Far Eastern Russia, September
Sidewinder / Peringueys adder (Bitis peringueyi) lying in wait, buried in sand ready to ambush prey, Namib Desert, Namibia
Red knot (Calidris canutus rogersi) incubating nest on a coastal gravel spit. Chukotka, Russia. June
Greater sage-grouse (Centrocercus urophasianus) male displaying on a lek in snow, Sublette County, Wyoming, USA. April
Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards
Common toads (Bufo bufo) in mating ball underwater, Belgium, March
Florida manatee (Trichechus manatus latirostrus) with Blue gill sunfish (Lepomis macrochirus) cleaning it, in a freshwater spring, beneath trees. Three Sisters Spring, Crystal River, Florida, USA
White-beaked dolphin (Lagenorhynchus albirostris) porpoising, Varangerfjorden, Finnmark, Norway. March
Carrion crow (Corvus corone corone) mobbing Lammergeier (Gypaetus barbatus) Spain, November
Black Labrador retriever portrait in barley field, Wiltshire, UK
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
Black blotched pufferfish (Diodon liturosus) inflated with water at night as defensive behaviour, Sipadan Island, Borneo, Malaysia
Leaf cutter ant (Atta sp) female worker carrying leaf to nest, with a smaller minor ant riding. The minors job is to protect her sister from parasitic flies. Costa Rica, December
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
Male Andean Cock-of-the-Rock (Rupicola peruvianus) displaying. Cock-of-the-Rock Lodge, Peru, August
African Elephant (Loxodonta africana) bulls in dominance fight, Mono Pools national Park, Zimbabwe, Southern Africa
Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using Starlight Camera technology without artificial lighting
Trumpeter Swans (Cygnus buccinator) in winter morning mist, showing aggression during courtship behaviour, Mississippi River, Minnesota, USA, February
Male Atlantic salmon {Salmo salar} leaping, migrating upstream to spawn, Perthshire, Scotland, UK, October
Maine Coon female cat, Serafin, in fierce defensive posture
British Shorthair Brown Spotted cat, Tiger Lily, reaching out
