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A signal blenny (Emblemaria hypacanthus) displays its showy dorsal fin as it extends out of its tube home. La Paz, Mexico. Sea Of Cortez, East Pacific Ocean
Male King eider (Somateria spectabilis) displaying, Norway, February
Osprey (Pandion haliaetus) in flight with a fish, Finland, July
RF - Polar Bear (Ursus maritimus) males fighting, Churchill, Canada, November
Lambs in meadow in spring, UK, April
Polar bear (Ursus maritimus) 3-year-olds play fighting on newly formed pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Pine marten (Martes martes) leaping, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Animal Behaviour category of the BWPA (British Wildlife Photography Awards) Competition 2016
Atlantic Puffins (Fratercula arctica) three resting on a snow bank, Hornoya, Varanger, Norway, April
Strawberry poison frog (Oophaga pumilio) pair mating, Sarapiqui, Heredia, Costa Rica
Pebas stubfoot toad / Harlequin toad (Atelopus spumarius) stretching on branch, Yasuni National Park, Orellana, Ecuador, Vulnerable Species
Black-headed bushmaster (Lachesis melanocephala) with tongue extended, Siquirres, Limon, Costa Rica
Scarlet macaws (Ara macao) on salt lick, Tambopata, Madre de Dios, Peru
Booted racket tail (Ocreatus underwoodii) two males fighting in flight, Mindo, Pichincha, Ecuador
Female Sea lion (Zalophus wollebaeki) with pup suckling in shallow water, Floreana Island, Galapagos
White-fronted nunbird (Monasa morphoeus) perched with lizard prey, Tambopata National Reserve, Madre de Dios, Peru
Crimson-rumped toucanet (Aulacorhynchus haematopygus) calling, Mashpi, Pichincha, Ecuador
Humpback Whale (Megaptera novaeangliae) breaching, Machalilla National Park, Manabi, Ecuador
Scarlet macaws (Ara macao) and Blue and yellow macaws eating clay close to the Tambopata river. Tambopata Reserve, Madre de Dios, Peru
Scarlet macaw (Ara macao) and Red and green macaw (Ara chloroptera) eating clay close to the Tambopata river, Tambopata Reserve, Madre de Dios, Peru
RF - Bottlenose dolphins (Tursiops truncatus) porpoising, Chanonry Point, Moray Firth, Highlands, Scotland. August
Lesser kestrel (Falco naumanni) male flying past during courtship with Mole cricket in its talons. Extremadura, Spain
Two male Lesser kestrels (Falco naumanni) in pursuit flight, one driving off the other from its nesting territory. Extremadura, Spain
Lioness (Panthera leo) chasing away Cheetah (Acinonyx jubatus) Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
RF - Head portrait of Spotted eagle owl (Bubo africanus), Kgalagadi Transfrontier Park, Northern Cape, South Africa, February 2016
Eagle owl (Bubo bubo) in flight through forest, backlit at dawn, Czech Republic, November. Captive
Little owl (Athene noctua) perched on a fence post, stretching its wings, Castro Verde, Alentejo, Portugal, April
RF- Eagle owl (Bubo bubo) calling, raised in captivity and newly released into the wild. Stockholm archipelago, Uppland, Sweden. September
Stellers Sea Eagles (Haliaeetus pelagicus) fighting on ice floe, Hokkaido, Japan. February
Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica
Fleischmanns glassfrog (Hyalinobatrachium fleischmanni) male attending eggs on the underside of a leaf overhanging a rainforest stream
Silk moth (Automeris zugana) sequence 2 of 2, with wings open to reveal eyespots, a means of deterring predators, photographed on a white background in mobile field studio
Four-spot orb weaver spider (Araneus quadratus) male approaching larger female to mate. Peak District National Park, Derbyshire, UK. October
Atlantic puffin (Fratercula arctica) gathering grass for its nest. Isle of Lunga, Treshnish Isles, Scotland, June
Emerald damselflies (Lestes sponsa) mating, with male above. Peak District National Park, Derbyshire, UK. September
Fly (Mycophaga testacea) laying eggs on Porcelain fungus (Oudemansiella mucida) growing on dead Beech tree (Fagus sylvatica). Peak District National Park, Derbyshire, UK. October
Bottlenose dolphin (Tursiops truncatus) porpoising, Moray Firth, Highlands, Scotland, UK. June
Grass snake (Natrix natrix) flicking tongue, Hampstead Heath, London, England, UK, August
Sand lizard (Lacerta agilis) male basking. Dorset, England, UK, May
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February
Red deer (Cervus elaphus) stag bellowing. Surrey, UK, October
Bank vole (Clethrionomys glareolus) feeding on blackberries, Worcestershire, England, UK, August. Meetyourneighbours.net project
Red kite (Milvus milvus) in a dive, Gigrin Farm, Rhayader, Wales, UK April
Kingfisher (Alcedo atthis) male taking off from water after diving for fish, Worcestershire, England, UK, July
Osprey (Pandion haliaetus) catching trout, Rothiemurchus estate, Cairngorms, Scotland, UK, July
Starling (Sturnus vulgaris) murmuration / mass gathering at winter roost, Salisbury Plain, Wiltshire, UK January
Robin (Erithacus rubecula) singing, Norfolk, England, UK, February
Pink anemonefish (Amphiprion perideraion) in Purple magnificent sea anemone (Heteractis magnifica) Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea