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Arctic Fox (Alopex / Vulpes lagopus) standing on ridge, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Arctic Fox (Alopex / Vulpes lagopus) yawning, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Japanese cranes (Grus japonensis) displaying in snow, Hokkiado, Japan, February
Lions (Panthera leo) - two brothers patrolling territorial boundary, affectionate behaviour, border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Polar bear (Ursus maritimus) juvenile shaking off snow after rolling around on the pack ice, arctic coast, 1002 area of the Arctic National Wildlife Refuge, Alaska, Beaufort Sea
Great crested grebe (Podiceps cristatus) carrying chick on back, The Netherlands, May
Sperm whale (Physeter macrocephalus) tail fluke above water as it dives below a rainbow, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species. Digital composite
Rainbow lorikeets (Trichoglossus moluccanus) pair preening each other, perched in tree. Ulladulla, New South Wales, Australia
Vogelkop superb bird-of-paradise (Lophorina niedda) male, performing courtship display on his display log, flashing false blue eye spots, a bright yellow gape, and blue breast shield
Green tree ants (Oecophylla smaragdina) defending their leaf nest, Daintree River, Wet Tropics World Heritage area, Queensland, Australia
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair close-up Manuel Antonio National Park, Quepos, Costa Rica
Bengal Tiger (Panthera tigris) six month old cub jumping on its mother, Bandhavgarh National Park, India
Andean cock-of-the-rock (Rupicola peruvianus), male displaying at lek in tree. Manu Biosphere Reserve, Amazonia, Peru
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement
Brown hare, (Lepus europaeus), group of animals in field, Islay, Scotland, UK. March
Great Spotted Woodpecker (Dendrocopos major) adult male feeding young, Oisterwijk, The Netherlands, June
Piping Plover (Charadrius melodus) brooding three chicks with a fourth approaching, northern Massachusetts, USA.June. Endangered species
Great egret (Ardea alba) two fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Tabby kitten, 6 weeks, lying with head up and raised paw
Badger (Meles meles) adult, backlit at sunset, scratching its ears, Derbyshire, UK, May. Winner of the Animal Behaviour category of the British Wildlife Photography Awards (BWPA) Competition 2017
RF- Western clownfish (Amphiprion oceallaris) in Magnificent sea anemone (Heteractis magnifica). Raja Ampat, West Papua, Indonesia
Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in march. Wapusk National Park. Churchill, Manitoba, Canada
Lion (Panthera leo) cubs playing, Masai-Mara Game Reserve, Kenya. Vulnerable species
Three Sperm whales (Physeter macrocephalus) socialising under the surface, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species
Trap-jaw ant (Odontomachus hastatus) with mandibles open, Los Amigos Biological Station, Peru
White-tailed eagle (Haliaeetus albicilla) taking off with fish prey, Norway, August
Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Wren (Troglodytes troglodytes) singing, early morning in spring, Broxwater, Cornwall, UK. April
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) crossing the Mara River. Northern Serengeti, Serengeti National Park, Tanzania
Goshawk (Accipiter gentilis) plucking a Willow Grouse (Lagopus lagopus) Liminka, Finland, March
Scottish wildcat (Felis silvestris grampia) captive, endemic to Scotland. September
RF - Australian Labradoodle, pouncing playfully
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Coconut / Veined octopus (Amphioctopus marginatus) hunts in the sand at night, while illuminated by a divers torch beam, Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Pacific Ocean
Marine iguana (Amblyrhynchus cristatus) on shore, Punta Moreno, Isabela Island, Galapagos
Red squirrel (Sciurus vulgaris) gnawing red deer antler for minerals, Cairngorms National Park, Highlands, Scotland, UK, June
King cobra (Ophiophagus hannah) in strike pose, Malaysia
Tree sparrow (Passer montanus) feeding a fledgling on a branch of Hawthorn (Crataegus monogyna) blossom, Perthshire, Scotland. June 2013
Backlit photo of a Robust ghost pipefish (Solenostomus cyanopterus) reveals a small fish that it has just eaten in its stomach. Lembeh Strait, North Sulawesi, Indonesia. Molluca Sea
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Caspian Tern (Sterna caspia) in flight, diving for prey. Lake Ellesmere, Canterbury, South Island, New Zealand
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Bottlenose dolphins (Tursiops truncatus) at the surface at sunrise. Kwazulu-Natal, South Africa
