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White-throated Mountain Gem (Lampornis castaneoventris), Green-crowned Brilliant (Heliodoxa jacula) group of three with one in flight, Savegre, Costa Rica
Wild Eurasian lynx (Lynx lynx) at night with city lights and sky glow behind, Switzerland. December. Taken with remote camera trap
Group of Great egrets (Ardea alba), Grey heron (Ardea cinerea) and Eurasian spoonbill (Platalea leucorodia) at night with stars and moon. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. May
Grass snake (Natrix natrix) in habitat, Aland Islands, Finland, May
Leaf cutter ants (Atta sp) carrying plant matter, Costa Rica
Solar eclipse, Finland, 20 March 2015
Downy emerald dragonfly (Cordulia aenea) in flight, Joutsa, Finland, July
Common raven (Corvus corax) two adults in flight, Hornoya, Norway, April
Hawfinch (Coccothraustes coccothraustes), male perched on twig, Uto, Finland, May
Silver washed fritillary butterfly (Argynnis paphia) female in flight with thistles, Uusimaa, Finland, August
Black-and-white kitten, Solo, 6 weeks, taking a selfie. Composite image
Northern gannets (Morus bassanus). Bass Rock, Scotland, UK. August
View up to Beech (Fagus sylvatica) woodland canopy in winter, photographed with a circular fisheye lens. Derbyshire, UK. December
Milky Way rising over the south wall of the Grand Canyon, Arizona, USA. November 1 2015. Composite image
European Chubb (Squalius / Leuciscus cephalus) in cave, Plitvice Lakes National Park, Croatia. November
Scorpion (Heterometrus sp) fluorescing in ultraviolet light at night on forest floor, Danum Valley, Sabah, Borneo
Pebbles reflected in the water surface at mouth of Sheryldy river, Lake Baikal, Siberia, Russia. September
Andromeda (M31) with its two syster galaxies (M32 and M110) Eastern Colorado. 8 November 2015. Taken with digital focus stacking
Underwater view of the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
European brown bear (Ursus arctos) head portrait, captive, occurs in Eurasia
Mute swan (Cygnus olor) flock on snow, Hazerswoude, The Netherlands, February
Golden eagle (Aquila chrysaetos) portrait, captive, occurs in the Northern hemisphere
Bealach na Ba Pass, Wester Ross, Scotland, UK, November 2015
Robin (Erithacus rubecula) profile in field studio, Scotland, UK, January
Black and white photograph of saltmarsh at Annat, Beinn Torridon, Scotland, UK, November
Grey wagtail (Motacilla cinerea) at waters edge, Gaoligong Mountain National Nature Reserve, Yunnan Province, October
Whooper swans (Cygnus cygnus) in misty hot springs, Lake Kussharo, Hokkaido, Japan, February
European hamster (Cricetus cricetus) male, in underground burrow, captive
Tsaranoro massif, Andringitra NP, Madagascar, October 2015
Red-backed shrike (Lanius collurio) adult male taking off, Lower Saxony, Germany, June. Sequence 2 of 3
Atlantic Puffin (Fratercula arctica) covered by mud from its partner digging out their nesting chamber / burrow, Skomer Island, Wales, UK, April
Namib sand snake (Psammophis namibensis) in sand dunes, Swakopmund, Erongo Region, Namibia
Orange anemonefish (Amphiprion sandaracinos) in its host Mertens carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea
Flame anthias (Pseudanthias ignitus) displaying to the more drab female (above) during courtship in the evening. Similan Islands, Thailand. Andaman Sea, Indian Ocean
Young California sea lion (Zalophus californianus) play in the canopy of a forest of Giant kelp (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. California, USA. North East Pacific Ocean
Great hammerhead shark (Sphyrna mokarran) female at night with Bar jacks (Caranx ruber), South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Wolf fish (Anarhichas lupus) portrait, Akuyeri, Iceland. North Atlantic Ocean
Lion (Panthera leo) male sharpening his claws on a tree at night, Sabi Sand Game Reserve, South Africa
Ol Doinyo Lengai, an active volcano held in high esteem and known locally as The Mountain of God by the Masai, with people and huts on the savannah below, Rift Valley, Tanzania
Puma (Puma concolor) in high altitude habitat of Torres del Paine National Park, Chile
Prayer plant (Calathea plowmanii) close up of leaf, TU Delft Botanical Garden, Netherlands, August
Aerial view of eroded banks and river with suspended load, caused by deforestation, central Madagascar
Alpine ibex (Capra ibex) adult male in deep snow on a ridge with young during snowfall, Gran Paradiso National Park, the Alps, Italy. January
European brown bear (Ursus arctos) scratching up against tree in snow, Finland, April
Black bear (Ursus americanus) male close up head portrait, Minnesota, USA, June
Coyote (Canis latrans) jumping in snow with its prey, Yellowstone National Park, Wyoming, USA, February
Head portrait of grey Andalusian stallion in dark barn, Northern France, Europe
Grey Arabian stallion running in desert dunes near Dubai, United Arab Emirates