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Pleiades or Seven Sisters (Messier 45 aka M45) in Taurus Constellation, taken from Eastern Colorado, USA. 8 November 2015
North American Bison (Bison bison) coated in frost standing on snow, Yellowstone National Park, Wyoming, USA, January
Ocelot (Leopardus pardalis) portrait. camera trap image, Nicoya Peninsula, Costa Rica
Leaf cutter ants (Atta sp) carrying colourful plant matter, reflected in water, Laguna del Lagarto, Santa Rita, Costa Rica
Kottigehara dancing frog (Micrixalus kottigeharensis), dancing frog name given because of foot flagging behavior between competing males. Endemic to Western Ghats. Agumbe, Karnataka, India
Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
Blue whale (Balaenoptera musculus) underwater, Coronado Islands, Baja California, Mexico, Eastern Pacific Ocean
Red fox (Vulpes vulpes) at night in snow, camera trap image, Jura Mountains, Switzerland, August. Commended in the Mammals category of the Asferico Competition 2016
Tree ferns (Cyatheaceae) in Kells Bay Gardens, Ring of Kerry, Iveragh Peninsula, County Kerry, Ireland, Europe. September 2015
Fiordland crested penguins (Eudyptes pachyrhynchus) in dense coastal forest, Lake Moeraki, South Island, New Zealand, November, threatened species
Common loon (Gavia immer) nesting at edge of lake and calling, Acadia National Park, Maine, USA, August
Aggregation of Golden jellyfish (Mastigias sp.) in a marine lake in Palau, the golden colour of this species comes from symbiotic algae in its tissues
Cow killer velvet ant (Dasymutilla occidentalis) on white background, Tuscaloosa County, Alabama, USA September
Whale shark (Rhincodon typus) viewed from above, Cenderawasih Bay, West Papua. Indonesia
Mexican free-tailed bats, (Tadarida brasiliensis), leaving Bracken Cave, Texas
Malabar torrent toad (Ghatophryne ornata) with bright yellow-orange spots more on abdomen. Coorg, Karnataka, India. Endemic to the Western Ghats. Endangered species. Endangered species
Patterdale terrier dog puppy, Korka, age 4 months
Fungal mycelium from an unidentified species of fungus, growing across a decomposing leaf. The mycelium is the vegetative part of the fungus and is made up of a mass of thread-like hyphae
Green iguana (Iguana iguana) rear view of basking in sun, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico, May
Bearded reedling (Panurus biarmicus) on frosty reed seed head, Helsinki, Finland, January
Thresher shark (Alopias pelagicus) swimming over seabed to be cleaned by cleaner wrasses, Cebu, Malaspascua, Philippines, September
Grey seal (Haichaoerus grypus) swimming below surface with sunbeams. Lundy Island, Devon, UK. Bristol Channel, August
Grass fire at night in Pantanal, Brazil
Dalmatian pelican (Pelecanus crispus) close up of feathers, Danube Delta, Romania
Bauxite mine, blue water caused by industrial runoff, minerals into local water, Linden town, Guyana, South America
Montane horned frog (Megophrys kobayashii) in the leaf litter at night. Mount Kinabalu (around 1400m asl), Sabah, Borneo
Lava flow from Kilauea Volcano flowing into the Pacific Ocean, Kalapana Coast, Big Island, Hawaii. August 2016
A Cotswold Lion sheep, rare breed, Gloucestershire, UK. June
Skeleton of a seahorse (Hippocampus sp. ) showing the remarkable structure of bonesSkeleton of a seahorse (Hippocampus sp.) showing the remarkable structure of bones that form a virtual exoskeleton, with few deep internal bones, in this unusual vertebrate group
Historical illustration of Common octopus (Octopus vulgaris) Pierre Denys de Montfort engraving from Histoire Naturelle Generale et Particuliere des Mollusques, 1801-1802
Aerial view of Giant water lily (Victoria amazonica) leaves in river, i Rurununi savanna, Guyana, South America
Grand Canyon at dusk, abstract hazy landscape, Arizona, USA, July
Moose (Alces americanus) close-up grazing, Denali National Park, Alaska, USA, September
Yellow warty frogfish (Antennarius maculatus) Seraya, Tulamben area, Bali, Indonesia. Java Sea
Mount Roraima tepui (flat top mountain) is the highest of the Pakaraima chain of Tepuis plateaus in South America, Pakarajma Mountains, near Phillipai, West Guyana, South America
Galapagos racer snakes (Pseudalsophis biserialis) group alert watching for prey on beach, Galapagos, June. Endangered species. Taken on location for BBC tv Blue Planet 2 series
Sable antelope (Hippotragus niger) bull charging through the shallows of the Selinda Spillway, with another behind, northern Botswana, August
European robin (Erithacus rubecula) low angle view near church, Bristol, UK. January
Wild Eurasian lynx (Lynx lynx) at night with city lights and sky glow behind, Switzerland. December. Taken with remote camera trap
Common eider (Somateria mollissima), female on her nest with sea behind, Finland, May
Low angle view of Porcelain fungus (Oudemansiella mucida) growing on a dead Beech tree (Fagus sylvatica). Plitvice Lakes National Park, Croatia. November
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
Black faced sheep ram with twisted horns, Mull, Scotland, UK. January
Zebra (Equus quagga) leaping during stampede, Serengeti, Tanzania. Vignette added and right edge expanded
Night-blooming cereus cactus (Acanthocereus tetragonus), flower bud opening at night, Texas, USA. Sequence 7 of 7. August
Polar bear (Ursus maritimus) spring cub sitting upright on hind legs balancing, Bernard Spit, 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Giant Manta (Manta birostris) with Remoras (Remora brachyptera) attached to both wings, offshore Santa Maria Island, Azores, Atlantic Ocean
Septarian concretion or (septarian nodules) from Morocco
Young Polar bear (Ursus maritimus) standing on hing legs, Bernard Spit, 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Siphonophore (Nanomia sp) close up of nectophores, and palpons. Gulf of Maine, Atlantic Ocean. Epipelagic species found up to 300m depth
Coconut / Veined octopus (Amphioctopus marginatus) hunts in the sand at night, while illuminated by a divers torch beam, Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Pacific Ocean
Bignose unicornfish (Naso vlamingii) profile of male displaying (the blue stripe at the bottom of the frame is the tail of another individual) Baa Atoll, Maldives. Indian Ocean
African wild dogs (Lycaon pictus) play fighting, Mkuze, South Africa
Three metre tall Mount Teide bugloss / Tower of jewels / Red Tajinaste (Echium wildpretii) flowering beside Teide wallflowers (Erysimum scoparium) on misty mountainside, Teide National Park
Indonesian blue-tongued skink (Tiliqua gigas) with tongue extended, captive, from Papua New Guinea
Cave Pearls, formed as dripping water rich in calcium salts deposits calcite around a small nucleus (often a sand grain)
Miniature American shepherd puppy in play-bow
Black smoke fluffy Lionhead x Lop rabbit
Slime mould (Physarum sp) plasmodium growing across rotting wood, Osa Peninsula, Costa Rica
Moss mimic stick insect (Trychopeplus laciniatus) showing amazing camouflage on mossy vine, photographed in tropical rainforest in mobile field studio against a white background
Strawberry poison frog (Oophaga pumilio) photographed on a white background in mobile field studio, Central Caribbean foothills, Costa Rica
Bearded tit (Panurus biarmicus) male portrait feeding, Helsinki, Finland, January
Green june beetle (Cotinis nitida) in flight Williamson County, Texas, USA Controlled conditions. October
Tabby kitten, 6 weeks, lying with head up and raised paw
Kelp forest (Laminaria hyperborea), Atlantic Ocean, North West Norway, March
Wild Salt River stallion grazing in water, living wild in Tonto National Forest, Arizona, USA. October
Red bat (Lasiurus borealis) female flying; shots taken with high speed flash San Saba County, Texas, USA. Controlled conditions. July
Blackbird (Turdus merula) in flight to feed on berries, Helsinki, Finland. November
Composite photograph showing diversity of colour and pattern in samples of granite
American crocodile (Crocodylus acutus) rear view of animal resting in shallow water with head at surface, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico
Ptarmigan (Lagopus mutus) in snowy landscape, Utsjoki, Finland, February
Part of a submerged Holocene forest, revealed after a winter storm from beneath the sand at Hells Mouth (Porth Neigwl), Abersoch Wales, UK March 2014
Great grey owl (Strix nebulosa) taking off from snowy ground, Kuusamo, Finland. March
Red fox (Vulpes vulpes) resting with eyes open, Vanoise National Park, Rhone Alpes, France. October
Shore of lake in Torres del Paine. Patagonia, Puerto Natales, Chile
Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean
Capercaillie (Tetrao urogallus) male displaying in rain, Kuusamo, Finland. March
Panoramic landscape of Cerredo mountain at sunset, Castro Urdiales, Cantabria, Spain
Eurasian / European badger (Meles meles) outside urban sett behind wall with graffiti, Bristol, UK May
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Ivy (Hedera helix) covered tree trunks in Knight Wood, Hampshire, UK, October
Common / Northern minke whale (Balaenoptera acutorostrata) surfacing, Pacific coast of Baja California, Mexico, February