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Altai argali sheep (Ovis ammon), three grazing on slope. Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China. October
Tibetan gazelle (Procapra picticaudata) in snow. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October
Face portrait of a Great hammerhead shark (Sphyrna mokarran) off Bimini, Bahamas
Eclectus parrot (Eclectus roratus) two males arrive at a nest cavity at the same time, both to provision the female within. Iron Range National Park, Cape York Peninsula, Queensland, Australia
Magnificent riflebird (Ptiloris magnificus alberti) male, on his display perch performing a display. Piccaninny Plains Sanctuary, Cape York Peninsula, Queensland, Australia
RF-Asian elephant (Elephas maximus) dust bathing. Jim Corbett National Park, Uttarakhand, India. (This image may be licensed either as rights managed or royalty free)
Pallass cat (Otocolobus manul) licking lips whilst running over ice. East Mongolia. February
Canopy jumping spider (Phidippus otiosus) male, North Florida, USA, September
Veined Octopus, Coconut Octopus (Amphioctopus marginatus, formerly Octopus marginatus), inhabiting discarded bivalve mollusc shells, which it collects and pulls around itself - defense, shelter
Snow leopard (Panthera uncia) with ears back. Captive
Eurasian lynx (Lynx lynx) yawning. Captive
Giant tun (Tonna galea) a species of marine gastropod mollusc that is one of the biggest sea snails in the Mediterranean, photographed off Vis Island, Croatia, Adriatic Sea
Hyacinth macaws (Anodorhynchus hyacinthinus) squabbling on palm tree, Pantanal, Mato Grosso, Brazil
Reef manta ray (Manta alfredi) filter feeding on plankton. Madhivafaru Reef, Raa Atoll, Maldives
Lion (Panthera leo) males mock fighting / play fighting. Ngorongoro Crater, Ngorongoro Conservation Area (NCA), Tanzania
Red fox (Vulpes vulpes) running through snow. Hayden Valley, Yellowstone, USA. January
Fosa / Fossa (Cryptoprocta ferox) male resting in dry decidous forest. Kirindy, western Madagascar. Endangered
Southern right whale (Eubalaena australis) diving, with tail fluke splash. Monumento Natural Ballena Franca Austral, UNESCO World Heritage Site, Valdes Peninsula, Patagonia, Argentina
Sunda clouded leopard (Neofelis diardi) at night, Deramakot Forest Reserve, Sabah, Malaysian Borneo
Common Loons (Gavia immer) one diving underwater, Michigan, USA. June
Moose bull (Alces alces) walking in forest clearing, Denali National Park, Alaska, USA, September
Black oil beetle (Meloe proscarabaeus) male on grass stem in sand dunes, Gower, Wales, UK, February. Focus stacked image
RF - Oceanic whitetip shark (Carcharhinus longimanus) swims in open waters, close to the surface with sun beams. Egypt. Red Sea. (This image may be licensed either as rights managed or royalty free.)
Javan leopard (Panthera pardus melas) roaring, native to the Indonesian island of Java. Captive. Digital composite
Porto Santo Wolfspider (Hogna schmitzi) Porto Santo island, Madeira Archipelago, Portugal. Endemic
Indri (Indri indri) with young, Palmarium Resrve Ankanin ny Nofy, Littoral rainforest, Madagascar
Great Grey Owl (Strix nebulosa) Kuhmo Finland, March
Sparrowhawk (Accipiter nisus) adult female, Norway, October
Golden jackal (Canis aureus) pup sitting on a grassy verge by a path, Danube Delta, Romania, July
African lion (Panthera leo) and lioness after mating, Masai Mara National Reserve, Kenya
Lion (Panthera leo) male walking. Masai Mara National Reserve, Kenya
Lady Amhersts pheasant (Chrysolophus amherstiae) male displaying, Kanding, Sichuan, China. April
European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition
Polar bears (Ursus maritimus) fighting in water. Beaufort Sea, Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Natures Best Photography Competition 2019
Grey-headed flying-fox (Pteropus poliocephalus) spotting and flaring wings to come in to land on a branch. Yarra Bend Park, Kew, Victoria, Australia. ÔÇ¿November 2019
Common Frog (Rana temporaria), and frogspawn in a pond, Coldharbour, Surrey, England. March. Highly commended in the Animal Portrait category of the BWPA Competition 2019
Hard coral (Porites lutea) with Christmas-tree worms (Spirobranchus giganteus), New Caledonia, Pacific Ocean
Bee-eater (Merops apiaster) male offering wasp as nuptial gift, Hungary. June
Common Cuckoo (Cuculus canorus) perched on gate Surrey, England, UK. April
Starling (Sturnus vulgaris) murmuration at sunset, West Pier, Brighton, England, UK
Timber rattlesnake (Crotalus horridus) females and newborn young at maternity site. Pennsylvania, USA. August
Fosa of Fossa (Cryptoprocta ferox) male walking through dry deciduous forest, Kirindy, western Madagascar
Aye-aye (Daubentonia madagascariensis) adult active and foraging in forest canopy at night. Deciduous forests, Daraina, northern Madagascar. Endangered endemic species
Red fox (Vulpes vulpes) adult hunting for rodents by snow diving in deep snow. Hayden Valley, Yellowstone, USA. January
Boky-boky (Mungotictis decemlineata) or Narrow-striped mongoose adult foraging in leaf litter. Dry deciduous forest, Kirindy, western Madagascar
Fosa or Fossa (Cryptoprocta ferox)(sometimes erroneously fossa) male in dry deciduous forest. Kirindy, western Madagascar
Puma (Puma concolor puma), female sitting in grassland. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. November
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
