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Rock formation caused by erosion in sandstone, Devils Garden, Grand Staircase-Escalante National Monument, Utah, USA, March 2014
American red squirrel (Tamiasciurus hudsonicus) collecting, Douglas Fir (Pseudotsuga menziesii) cones, Montana, USA. October
Yellow warbler (Dendroica petechia) collecting nesting material from Bulrush cattail (Typha sp) Bozeman, Montana. USA, June
Grizzly Bears (Ursus arctos) standing in heavy rain, Lake Clarke National Park, Alaska, September
Grizzly Bears (Ursus arctos) resting, Lake Clarke National Park, Alaska, September
Pika (Ochotona princeps) gathering plants to store for winter, Bridger National Forest, Wyoming, USA. August
RF -Three Bison (Bison bison) walking through snow with herd feeding in background. Yellowstone National Park, Wyoming, USA. January
Coyote (Canis latrans) in snow, Yellowstone National Park, Wyoming, USA, February
Grizzly cubs (Ursus arctos) two play fighting in the mist, near Brooks Falls in Katmai National Park, Alaska, USA, July
Grizzly Bear (Ursus arctos) resting, Lake Clarke National Park, Alaska, September
California sea lions (Zalophus californianus) playing in a kelp forest off Santa Barbara Island, California, USA
RF - Coyote (Canis latrans) in snow, Yellowstone. February
Red fox (Vulpes vulpes) resting and sitting on snow, Yellowstone National Park, USA, February
Coastal brown bear (Ursus arctos), Lake Clarke National Park, Alaska, September
Brown Bear (Ursus arctos) mother and 3-4 month cubs foraging for clams. Katmai National Park, Alaska, USA, June
Black bear (Ursus americanus) mother with cub. Yellowstone National Park, Wyoming, USA, May
Coastal brown bear (Ursus arctos) fishing, Lake Clarke National Park, Alaska, September
Porcupine (Erethizon dorsatum) on tundra. North Slope, Alaska, USA. September
Coyote (Canis latrans) in wintry landscape, Yellowstone National Park, USA, February
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Bison (Bison bison) standing in geothermal run-off in winter, Yellowstone National Park, Wyoming, USA, February
Sea otter (Enhydra lutris) floating on its back at surface among kelp, Alaska, USA. Gulf of Alaska, Pacific ocean
Bighorn Sheep (Ovis canadensis) on the hillside, ruminating, Yellowstone National Park, Wyoming, USA. January 2011
Coyote (Canis latrans) concealed in the snow, Yellowstone National Park, Wyoming, USA. January
Red fox (Vulpes vulpes) in snow, Yellowstone National Park, Wyoming, USA. Wyoming, USA, January
Bison (Bison bison) standing steam from geothermal springs, Yellowstone National Park, Wyoming, USA, February
RF- Bison (Bison bison) standing in steam from geothermal springs. Yellowstone National Park, Wyoming, USA, February
Polar bear (Ursus maritimus) sow with a pair of cubs walk on a barrier island during autumn freeze up, Bernard Spit, North Slope, Arctic coast of Alaska, September
Yellow warbler (Dendroica petechia) male singing. Bozeman, Montana, USA
Bald eagle (Haliaeetus leucocephalus) in flight, Homer, Alaska, USA, March
RF - Great horned owl (Bubo virginianus) in woodland, captive; Alaska, USA
Short-billed Dowitcher (Limnodromus griseus) on mudflats. Grays Harbor County, Washington. May
Barred Owl (Strix varia) fledgling resting in tree, King County, Washington, USA May
Spotted Owl (Strix occidentalis). Willamette National Forest, Oregon. June
Spruce grouse (Dendragapus fuliginosus) stops while foraging among fall blueberry bushes. Denali National Park, Alaska, USA
Horned puffins (Fratercula corniculata) pair interacting by touching bills while perched on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Lazuli Bunting (Passerina amoena) perched, Grand Teton National Park, Wyoming, USA, June
Burrowing owl (Athene cunicularia) Adams County, Washington, USA. May
Lower Falls at Artist Point, Yellowstone River. Yellowstone National Park, Wyoming, USA, June
Polar bear (Ursus maritimus) sow with two juveniles walk along the ice edge during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Grey wolf (Canis lupus) Katmai National Park, Alaska, USA, August
Rock formations silhouetted at sunset on the Pacfic coast of Olympic National Park, Washington State, USA
Wild Mustang pinto foal nuzzling up to mother, Sand Wash Basin Herd Area, Colorado, USA
Wild Mustang foals among wild flowers, Pryor Mountains, Montana, USA
Water rushing down alpine stream, Logan Pass, Glacier National Park, Rocky Mountains, Montana, July 2010
Grey wolf (Canis lupus) portrait, Katmai National Park, Alaska, USA, August
Calcified trees at Tangled Creek in winter, Yellowstone National Park, Wyoming, USA, February 2013
Sandstone cliff at sunset, Colorado Plateau, Kodachrome Basin State Park, Utah, USA November 2012
