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Young Polar bear (Ursus maritimus) standing and trying to balance in shallow water along the Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska
Mountain Gorilla (Gorilla beringei beringei) Sabyinyo Group, Silverback in meadow, Volcanoes National Park, Rwanda, East Africa
Ghost glass frog (Centrolenella ilex) portrait, Costa Rica
Golden retriever sitting on boat beside water, USA
Goldfinch (Carduelis carduelis) amongst Blackthorn blossom, Hertfordshire, England, UK
European grey wolf (Canis lupus) interacting with a European brown bear (Ursus arctos) Kuhmo, Finland, July 2009. WWE INDOOR EXHIBITION
Barn owl (Tyto alba) in flight, Norfolk, England, UK, February
Ural Owl (Stix uralensis) resting in snowy tree, Kuusamo Finland February
Mountain hare (Lepus timidus) silhouette at dusk in late summer. Scotland, October
Gannet (Morus bassanus) adult turns in the air as it begins to plunge dive. Shetland Islands, Scotland, UK, July
Common Kingfisher (Alcedo atthis) adult male at rest, Dorset, UK, November
Two Bewick swans {Cygnus columbianus} flying with full-moon in sky. UK
Portrait of a young Little Owl (Athene noctua)
Blue footed booby displaying {Sula nebouxii} Galapagos
Masai giraffe {Giraffa camelopardalis} mother nuzzling baby, Lower Masai Mara GR, Kenya
Bottlenose dolphin (Tursiops truncatus) two breaching in evening light, Moray Firth, Inverness-shire, Scotland, UK, August, sequence 2/2
Duckling swimming on water surface, split level, captive, UK
Snowy Owl (Nyctea scandiaca) adult female flying over snow, winter, Europe Captive /Snowy Owl (Nyctea scandiaca) adult female flying over snow, winter, Europe Captive/ Falconers bird, Austria
Adelie penguins (Pygoscelis adeliae) diving off iceberg, Antarctica, January
Two white horses of the Camargue, head to head at water, Camargue, Southern France
Griffon vulture (Gyps fulvus) in flight, Monfrague National Park, Extremadura, Spain, March 2009
Red kite (Milvus milvus) in flight, Gigrin Farm, Mid Wales, UK, March. Non-ex
Dipper (Cinclus cinclus) singing from prominent rock in breeding season, Derbyshire, UK
Curlews (Numenius arquata) group flying over the sea during storm. Wales, UK December. Highly commended, Habitat division, British Wildlife Photography Awards (BWPA) competition 2012
Portrait of English Springer Spaniel (field type). Elkhorn, Wisconsin, USA, January
Wandering albatross {Diomedea exulans} flying over open ocean, South Atlantic
Asian Elephant (Elephas maximus) male, India
Asian Elephant (Elephas maximus) male, walking through forest, South India
Golden Retriever puppy in basket with flowers. USA
Head portrait of a Giant panda (Ailuropoda Melanoleuca) Bifengxia Giant Panda Breeding and Conservation Center, Yaan, Sichuan, China
Blacktail Prairie Dog (Cynomys ludovicianus) engaging in Jump-yip behavior - A strong arch of the back or " jump" followed by a shrill " yip" - Thought to occur when a predator
Common opossum {Didelphis marsupialis} playing dead as defence strategy
Cavalier King Charles Spaniel, puppy, black-and-tan, 6 weeks, sitting on grass, wearing collar
Platypus {Ornithorhynchus anatinus} swimming underwater Tasmania, Australia. Commposite
Seven Ravens (Corvus corax) perched on dead tree in the snow, Utajarvi, Finland, February
Inflated spotted porcupinefish (Diodon hystrix), Maui, Hawaii
Three Whooper swans (Cygnus cygnus) in flight, Lake Tysslingen, Sweden, March 2009 WWE BOOK. WWE INDOOR EXHIBITION
Portrait of Ballan wrasse (Labrus bergylta), St Abbs (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, UK, October 2011
Tricolour Border Collie puppy
Adelie penguin (Pygoscelis Adeliae) in falling snow on the western Antarctic Peninsula, Southern Ocean
Bittern (Botaurus stellaris) in flight, Minsmere RSPB reserve, Suffolk, UK, August
Bat fish juvenile {Halicutaea sp} deep sea
Starry smooth-hound shark (Mustelus asterias) resting on seabed, Channel Isles, UK
Jaguar (Panthera onca) close-up head portrait, lying down, captive
Three Red Kites (Milvus milvus) chasing each other in flight. Wales, UK. November
Yellow-headed Picathartes / White necked rockfowl (Picathartes gymnocephalus) Kambui Hills, Sierra Leone, Vulnerable species. Photograph taken on location for BBC Africa series, September 2010
Philippine or Monkey eating eagle, Philippines (captive)
Atlantic Puffin (Fratercula artica) adult on breeding cliffs. Hermaness NNR, Shetland, UK, June