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Wolf (Canis lupus) pack running in deep snow, captive in enclosure of the Bavarian Forest National Park, Germany, February
Whale Shark (Rhincodon typus) passing by baitball, Santa Maria Island, Azores, Portugal, September 2008 Exclusive Japanese calendar rights for 2014
Portrait of a grey Andalusian stallion, in Ciutadella, Menorca, Spain
Atlantic Sailfish (Istiophorus albicans) attacking school of sardine (Sardinella aurita) bait ball. Isla Mujeres, Caribbean Sea, Mexico
Female Scottish wildcat (Felis silvestris grampia) portrait, in winter coat, semi habituated living wild in native oakwood, Lochaber, Scotland, April
Atlantic puffin (Fratercula arctica) gathering around burrows on cliff top, Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland, UK, June
Adult Adder (Vipera berus) basking on pebble beach. Abbotsbury, Dorset, UK, March
Suffolk Punch heavy horse in field resting head anothers back, UK, September
Norfolk terrier portrait, lying in autumn foliage
Silhouette of Barn Swallow (Hirundo rustica) flying over water, hawking for insects, Berwickshire, Scotland, UK, July
Eastern Grey Bamboo Lemur (Hapalemur griseus) feeding on bamboo shoot, Andasibe-Mantadia National Park, Madagascar, October
European Lynx (Lynx lynx) adult male stalking through winter birch forest, Bardu, Norway, captive
Male Sage Grouse {Centrocercus urphasianus} courtship display, Baggs, Wyoming, USA
Sea Otter floating at sea surface holding pup. Alaska (Enhydra lutris) USA
Impala with Oxpeckers. Kruger National Park, South Africa
Emperor penguins (Aptenodytes forsteri) explode out of the water, returning to breed at Cape Washington, Antarctica. Taken on location for BBC Frozen Planet series, 2009
Young Polar bear (Ursus maritimus) standing and trying to balance in shallow water along the Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska
Sea otter (Enhydra lutris) resting at the sea surface with paws in the air. Barkley Sound, Vancouver Island, Canada, August
Seven Little Bee-eaters (Merops pusillus) in a row, one facing away, Masai Mara, Kenya
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Red Fox (Vulpes vulpes) walking along a fallen trunk in misty forest. Black Forest, Germany, November
Southern cassowary (Casuarius casuarius), adult male with chicks keeping a wary eye on the photographer. Near Bloomfield, north Queensland, Australia, September
American white & Brown pelican P. occidentalis}, Florida, USA
Musk lorikeet (Glossopsitta concinna) feeding on flowers in a Eucalyptus tree, south-east Tasmania, Australia, February
White chinned petrel (Procellaria aequinoctialis) in flight low over the water off Kaikoura, Canterbury, New Zealand
Red Deer stag and hind (Cervus elaphus) silhouetted at sunset, Holkham Park, Norfolk, UK, December
Peregrine Falcon (Falco peregrinus) perched on a cliff above a woodland in autumn. Kinnoull Hill Woodland Park, Perthshire, Scotland, November
Gannet (Morus bassanus) fluffy chick with its parent. Shetland Islands, Scotland, UK, August
Three North American / Canadian Otters (Lutra canadensis) lying on each other by water. Captive, UK
Minature Shetland Pony in snow, UK
Laughing kookaburra (Dacelo novaeguineae) Tasmania, Australia
Lazuli Bunting (Passerina amoena) male in breeding plumage, singing, Mono Lake Basin, California, USA. July
Stac Lee and Stac an Armin, home to Northern gannet (Morus bassanus) colonies, St. Kilda Archipielago, Outer Hebrides, Scotland, UK, June 2009
Thrush nightingale (Luscinia luscinia) in tree singing, Matsalu National Park, Estonia, May 2009
Great grey owl (Strix nebulosa) close-up of head, Northern Oulu, Finland, June 2008
Brown trout (Salmo trutta) in shallow water migrating upstream, Bornholm, Denmark, November 2008
Common dolphins (Delphinus delphis) surfacing, Fisheye lens. Pico, Azores, Portugal, June 2009
European bison (Bison bonasus) behind tree, Bialowieza NP, Poland, February
Puffin (Fraterclua arctica) with fish in beak, Saltee Island, Ireland June 2009
Hoopoe (Upupa epops) landing on branch, Hortobagy NP, Hungary
European Bee-eater (Merops apiaster) pair, male passing Dragonfly prey to female, Pusztaszer, Hungary, May 2008
Blackbird (Turdus merula) male, portrait, Pusztaszer, Hungary, May 2008
Flock of Grey herons (Ardea cinerea) standing in a line on the edge of a pond, Regents Park, London, England, UK, May. Did you know
Lapwings (Vanellus vanellus), Golden plover (Pluvialis apricaria), Wigeon (Anas penelope) and Common Teal (Anas crecca) flying over flooded marshes in winter, with a rainbow in the background
Flock of Manx shearwaters (Puffinus puffinus) in flight over sea, near Ardnamurchan Point, Inner Hebrides, Scotland, UK, July 2011
Hen harrier (Circus cyaneus) adult female landing on rock in moorland habitat, Glen Tanar Estate, Grampian, Scotland, UK, June
Pike (Esox lucius) in disused quarry, Stoney Stanton, Stoney Cove, Leicestershire, UK, June
RF- Mute swan (Cygnus olor) on water in winter dawn mist. Loch Insh, Cairngorms National Park, Highlands, Scotland UK, December