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Wild boar (Sus scrofa) in old mixed conifer and broadleaf forest, Punia Forest Reserve, Lithuania, May
Smallspotted catshark (Scyliorhinus canicula) on sea floor, Jersey, British Channel Islands
Gorgonian / Sea whip coral (Ellisella ceratophyta) with Ring tailed cardinalfish (Apogon aureus) swimming in front, West Papua, Indonesia
Coral reef with soft corals (Dendronephthya sp. ), encrusting sponges and Pygmy sweepersCoral reef with soft corals (Dendronephthya sp.), encrusting sponges and Pygmy sweepers (Parapriacanthus ransonetti) Similan Islands, Andaman Sea, Thailand
European roller (Coracias garrulus) peering out of nest with feather, Lake Csaj, Pusztaszer, Kiskunsagi National Park, Hungary, February
Northern raven (Corvus corax) pair perching on branch. Danube Delta, Romania, May
Golden jackal (Canis aureus) portrait. Danube Delta, Romania, May
Gurskys spectral tarsier (Tarsius spectrumgurskyae) North Sulawesi
White tailed sea eagle (Haliaeetus albicilla) perched on tree stump with fish. Hooded crow (Corvus cornix) flying in to attack in background. Danube Delta, Romania. May
Horsfields tarsier / Western tarsier ( Tarsius bancanus ssp. saltator) Belitung Island, Sumatra, Indonesia. Endemic
South Georgia pintail (Anas georgica georgica) swimming in front of Southern elephant seal (Mirounga leonina), male. Gold Harbour, South Georgia. October
African wild dog (Lycaon pictus) portrait, Mana Pools National Park, Zimbabwe
Elephant (Loxodonta africana), male standing on hind legs to reach acacia pods with Cattle egrets (Bubulcus ibis) in background. Mana Pools National Park, Zimbabwe
Bee-eater (Merops apiaster) with Bumblebee prey, Sierra de Grazalema Natural Park, southern Spain, July
Iberian lynx (Lynx pardinus) with Magpie (Pica pica) behind, Sierra de Andujar Natural Park, Jaen, Spain, September
Grey whale (Eschrichtius robustus) eye, Magdalena Bay, Baja California, Mexico, February. Winner in the Ocean Views category of the 2018 Natures Best Winland Smith Rice Awards
Red fox (Vulpes vulpes) adult with train in background, Kent, UK. March. Highly commended in the Nature Photographer of the Year Awards 2018. Taken with remote camera
White rhinoceros (Ceratotherium simum) five calves, orphaned from poaching feeding from a trough at dusk at the Rhino Revolution orphanage near Hoedspruit, South Africa. May 2017
Great white egret (Ardea alba) group of three in winter, Kiskunsag National Park, Hungary. Winner of the Bird Category of Nature Photographer of the Year Awards 2018
Scarred face of a white rhinoceros (Ceratotherum simum) that survived an attack by poachers who illegally tranquillised her, two other rhinoceroses died from their injures
Aerial view of a herd of wild White rhinoceros (Ceratotherium simum) running free on Chiefs Island, Okavango Delta, Botswana
Young White rhinoceros (Ceratotherium simum) in a reinforced steel boma, blindfolded and partially drugged after a long journey from South Africa, before being released into the wild in Botswana
White-tailed eagle (Haliaeetus albicilla) male and female perched on tree in snow, near Kuusamo, Finland. March. Runner up in the Birds category of the GDT European Wildlife Photographer of the Year
Lava lizard (Microlophus peruvianus) juvenile peering out of eye of dead sea lion. Paracas National Reserve, Peru. Highly commended in the Animals in their Environment Category of Wildlife
Mountain hare (Lepus timidus) in extreme winds with spin drifts of snow, Cairngorms National Park, Scotland, UK, February 2018
Pallass cat (Otocolobus manul) Tibetan Plateau, Qinghai, China
Leopard (Panthera pardus) cub looking up at birds (out of frame) with mother in background, Khwai River, Botswana
Black spider monkey (Ateles chamek) calling, captive, Peru
White lipped peccary (Tayassu pecari) on forest floor, Lago Preto Conservation Concession, Yavari River, Peru
Black spider monkey (Ateles chamek) portrait, captive, Peru
Golden-mantled tamarin (Saguinus tripartitus) captive
Drill monkey (Mandrillus leucophaeus) adult male, portrait, captive
Great crested newt (Triturus cristatus) swimming
Victoria crowned pigeon (Goura victoria) head portrait, vulnerable species, captive
Shag (Phalacrocorax aristotelis) in flight over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Cameroon bush viper (Atheris broadleyi) captive occurs in West Africa
Beautiful pit viper (Trimeresurus venustus) captive occurs in Thailand
Razorbill (Alca torda) landing, Great Saltee Island, County Wexford, Ireland, June
Red-eyed tree frog {Agalychnis callidryas} resting in Bromeliad flower, Nicaragua, June
Puffin (Fratercula arctica) wings spread backlit, Great Saltee Island, County Wexford, Republic of Ireland, June
Ghost glass frog (Sachatamia ilex), Costa Rica
Snowy owl (Bubo scandiacus) in flight, Ontario, Canada, January
Wild Buffons macaw x Scarlet macaw (Ara ambiguus x Ara macao) hybrid, Costa Rica
Tibetan macaque (Macaca thibetana) Tangjiahe Nature Reserve, Sichuan Province, China
Chinese cobra (Naja atra) in threat stance, Shek Pik, southwestern coast of Lantau Island, Hong Kong, China
Indo-Pacific humpback dolphin (Sousa chinensis) with fishing net damage around throat, Pearl River Estuary, Lantau Island, Hong Kong, China
Thread-tail dartgoby (Ptereleotris hanae) Sharp Island or Kiu Tsui Chau, Kiu Tsui Country Park, Hong Kong, China
Fairy pitta (Pitta nympha) with a maggot in its beak, Guangshui, Hubei province, China. July
