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Common swift (Apus apus) group flying overhead, Bradford-on-Avon, Wiltshire, UK, May
Common European toad tadpoles (Bufo bufo), sequence showing development of limbs. Derbyshire, UK. August. Digital composite
White-throated magpie-jay (Calocitta formosa) Nicoya peninsula, Costa Rica. February
Adult Eyelash Pit Viper or Eyelash Viper (Bothriechis schlegelii) (Viperidae: Crotalinae). Distictive yellow / orange oropel form
Adult Aye-aye (Daubentonia madagascariensis) active in forest canopy at night. Dry deciduous forest near Andranotsimaty. Daraina, northern Madagascar. Critically Endangered
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Canebreak treefrog (Aplastodiscus sp) Guainumbi Private Reserve, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Eurasian hoopoe (Upupa epops) at nest in stone wall with hungry chicks gaping, Vendee, France, June
Clarion Island Whip Snake (Masticophis anthonyi), IUCN Critically Endangered, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage
Grey wolf (Canis lupus), adult with cub, captive, USA
Grass snake(Natrix natrix) coiled round eggs, Alsace, France
Scarlet ibis (Eudocimus ruber), two birds perched on bushes, Coro, Venezuela. Small repro only
Scarlet ibis (Eudocimus ruber), in flight, Coro, Venezuela
Nurse shark (Ginglymostoma cirratum) portrait, resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas. The Bahamas National Shark Sanctuary
Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
Southern Viscacha (Lagidium viscacia) at rest, Isla Incahuasi in the Salar de Uyuni, altiplano, Bolivia
Great Frigatebird (Fregata minor) arriving at nest with chick, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Zebra (Equus quagga) leaping during stampede, Serengeti, Tanzania. Vignette added and right edge expanded
Emerald toucanet (Aulacorhynchus prasinus) Rio Platano Biosphere Reserve and UNESCO World Heritage Site, La Mosquitia, Honduras
Nile crocodile (Crocodylus niloticus) Chobe National Park, Botswana, June
Galapagos sea lion (Zalophus wollebaeki) pup on shore, Galapagos
Giant panda (Ailuropoda melanoleuca) female, Huan Huan, holding baby age three months, Beauval Zoo, France, November 2017
Fin whale (Balaenoptera physalus) just below surface, south Barcelona coast, Spain, Mediterranean Sea
Whale headed / Shoebill (Balaeniceps rex) vocalising. Swamps of Mabamba, Lake Victoria, Uganda
Fin whale (Balaenoptera physalus) with ctenophore in front of its mouth, south Barcelona coast, Spain, Mediterranean Sea
Short-finned pilot whale (Globicephala macrorhynchus) showing underside whilst vertical underwater, South Tenerife, Canary Islands, Atlantic, October
Thomass Langur (Presbytis thomasi), animal endemic to northern Sumatra. These primates have relatively long arms and legs, adapted for an acrobatic life in the crowns of trees
Bairds tapir (Tapirus bairdii) Corcovado National Park, Costa Rica, May. Endangered
Stellers Sea Eagle (Haliaeetus pelagicus) with prey in beak in mid-air, Hokkaido, Japan, February
Close up if a Great hammerhead shark (Sphyrna mokarran) swimming over sandy seabed, South Bimini, Bahamas. The Bahamas National Shark Sanctuary, West Atlantic Ocean
Hoffmanns Two-toed sloth (Choloepus hoffmanni) mother and baby, aged 2 months, in tree, Costa Rica. Rescued and released by Aviarios Sloth Sanctuary
Proboscis monkey (Nasalis larvatus) mother suckling infant, Sabah, Malaysia
Harpy eagle (Harpia harpyja) close up head portrait, captive
Lammergeier / Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Song Thrush (Turdus philomelos), Bayern, Germany. June. June
Malabar torrent toad (Ghatophryne ornata) with bright yellow-orange spots more on abdomen. Coorg, Karnataka, India. Endemic to the Western Ghats. Endangered species. Endangered species
Giant Manta (Manta birostris) with Remoras (Remora brachyptera) attached to both wings, offshore Santa Maria Island, Azores, Atlantic Ocean
Young Polar bear (Ursus maritimus) standing on hing legs, Bernard Spit, 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Deccan ground gecko (Cyrtodactylus albofasciatus) on forest floor, backlit with water droplets. Amboli, Maharashtra, India. Endemic to Western Ghats
African wild dogs (Lycaon pictus) play fighting, Mkuze, South Africa
Group of four Giraffes (Giraffa camelopardalis) Mkuze, South Africa
Indonesian blue-tongued skink (Tiliqua gigas) with tongue extended, captive, from Papua New Guinea
Strawberry poison frog (Oophaga pumilio) photographed on a white background in mobile field studio, Central Caribbean foothills, Costa Rica
Wolf-eel (Anarrhichthys ocellatus) Vancouver Island, British Columbia, Pacific Ocean
Humpback whale (Megaptera novaeangliae) Umbili, a female age 8 years, breaching, Francisco Coloane Marine Park, Chiles first marine park, southeast of the Carlos III Island, Strait of Magellan, Chile
North Atlantic right whale (Eubalaena glacialis) tail fluke, Bay of Fundy, Canada
Killer whale (Orcinus orca) at surface, Johnstone strait, British Columbia, Canada
Pilot whales (Globicephala macrorhynchus) just below surface, Tenerife, Canary Islands, Atlantic Ocean
