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Melanistic leopard / Black panther (Panthera pardus) Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India
Desert hare (Lepus tibetanus)face and ears portrait, Inner Mongolia, China
Lemon shark pup (Negaprion brevirostris) in mangrove forest which acts as a nursery for juveniles of this species. Eleuthera, Bahamas
Lemon shark (Negaprion brevirostris) adult resting and allowing a small cleaner wrasse feed off parasites. Grand Bahama Island, Bahamas
Tapanuli orangutan (Pongo tapanuliensis) Togus, adult flanged male, Batang Toru Forest. Sumatran Orangutan Conservation Project, North Sumatran Province, Indonesia
Close view of a Green anaconda, (Eunectes murinus) underwater, Formoso River, Bonito, Mato Grosso do Sul, Brazil
Southern anteater (Tamandua tetradactyla) Formoso River, Bonito, Mato Grosso do Sul, Brazil
Red bat (Lasiurus borealis) female flying; shots taken with high speed flash San Saba County, Texas, USA. Controlled conditions. July
Blackbird (Turdus merula) in flight to feed on berries, Helsinki, Finland. November
Green iguana (Iguana iguana) rear view of basking in sun, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico, May
American crocodile (Crocodylus acutus) rear view of animal resting in shallow water with head at surface, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico
Bearded reedling (Panurus biarmicus) on frosty reed seed head, Helsinki, Finland, January
Great grey owl (Strix nebulosa) taking off from snowy ground, Kuusamo, Finland. March
Thresher shark (Alopias pelagicus) swimming over seabed to be cleaned by cleaner wrasses, Cebu, Malaspascua, Philippines, September
Red fox (Vulpes vulpes) resting with eyes open, Vanoise National Park, Rhone Alpes, France. October
Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean
Grey seal (Haichaoerus grypus) swimming below surface with sunbeams. Lundy Island, Devon, UK. Bristol Channel, August
Capercaillie (Tetrao urogallus) male displaying in rain, Kuusamo, Finland. March
Red-legged honeycreeper (Cyanerpes cyaneus) two males on barbed wire. Costa Rica
Flock of Tree sparrows (Passer montanus) in flight, with a Greenfinch (Chloris chloris) blurred motion, Pacsmag, Hungary
Nurse shark (Ginglymostoma cirratum) and Horse-eye jacks (Caranx latus) Shark Ray Alley, Hol Chan Marine Reserve, Belize Barrier Reef, Belize
Whale shark (Rhincodon typus) viewed from above, Cenderawasih Bay, West Papua. Indonesia
Skeleton of a seahorse (Hippocampus sp. ) showing the remarkable structure of bonesSkeleton of a seahorse (Hippocampus sp.) showing the remarkable structure of bones that form a virtual exoskeleton, with few deep internal bones, in this unusual vertebrate group
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Male Golden Pheasant (Chrysolophus pictus) standing on grass in display plumage. Kew Gardens, England, UK. March
Crocodile monitor (Varanus salvadorii) close up eye, captive, occurs in New Guinea
Pine grosbeak (Pinicola enucleator), male feeding juvenile, Finland, July
Black grouse (Lyrurus tetrix), male at lek, Finland, April
Long-tailed duck (Clangula hyemalis), males in flight, Finland, April
Kingfisher (Alcedo atthis) male, after diving, taking off from water with fish, a Common Roach (Rutilus rutilus) Lorraine, France, August
Common swift (Apus apus) group flying overhead, Bradford-on-Avon, Wiltshire, UK, May
Common European toad tadpoles (Bufo bufo), sequence showing development of limbs. Derbyshire, UK. August. Digital composite
White-throated magpie-jay (Calocitta formosa) Nicoya peninsula, Costa Rica. February
Adult Eyelash Pit Viper or Eyelash Viper (Bothriechis schlegelii) (Viperidae: Crotalinae). Distictive yellow / orange oropel form
Adult Aye-aye (Daubentonia madagascariensis) active in forest canopy at night. Dry deciduous forest near Andranotsimaty. Daraina, northern Madagascar. Critically Endangered
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Canebreak treefrog (Aplastodiscus sp) Guainumbi Private Reserve, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Eurasian hoopoe (Upupa epops) at nest in stone wall with hungry chicks gaping, Vendee, France, June
Clarion Island Whip Snake (Masticophis anthonyi), IUCN Critically Endangered, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage
Grey wolf (Canis lupus), adult with cub, captive, USA
Grass snake(Natrix natrix) coiled round eggs, Alsace, France
Scarlet ibis (Eudocimus ruber), two birds perched on bushes, Coro, Venezuela. Small repro only
Scarlet ibis (Eudocimus ruber), in flight, Coro, Venezuela
Nurse shark (Ginglymostoma cirratum) portrait, resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas. The Bahamas National Shark Sanctuary
Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
Southern Viscacha (Lagidium viscacia) at rest, Isla Incahuasi in the Salar de Uyuni, altiplano, Bolivia
Zebra (Equus quagga) leaping during stampede, Serengeti, Tanzania. Vignette added and right edge expanded
Emerald toucanet (Aulacorhynchus prasinus) Rio Platano Biosphere Reserve and UNESCO World Heritage Site, La Mosquitia, Honduras