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Great grey owl (Strix nebulosa) taking off from snowy ground, Kuusamo, Finland. March
Thresher shark (Alopias pelagicus) swimming over seabed to be cleaned by cleaner wrasses, Cebu, Malaspascua, Philippines, September
Red fox (Vulpes vulpes) resting with eyes open, Vanoise National Park, Rhone Alpes, France. October
Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean
Grey seal (Haichaoerus grypus) swimming below surface with sunbeams. Lundy Island, Devon, UK. Bristol Channel, August
Red-legged honeycreeper (Cyanerpes cyaneus) two males on barbed wire. Costa Rica
Flock of Tree sparrows (Passer montanus) in flight, with a Greenfinch (Chloris chloris) blurred motion, Pacsmag, Hungary
Nurse shark (Ginglymostoma cirratum) and Horse-eye jacks (Caranx latus) Shark Ray Alley, Hol Chan Marine Reserve, Belize Barrier Reef, Belize
Whale shark (Rhincodon typus) viewed from above, Cenderawasih Bay, West Papua. Indonesia
Skeleton of a seahorse (Hippocampus sp. ) showing the remarkable structure of bonesSkeleton of a seahorse (Hippocampus sp.) showing the remarkable structure of bones that form a virtual exoskeleton, with few deep internal bones, in this unusual vertebrate group
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Male Golden Pheasant (Chrysolophus pictus) standing on grass in display plumage. Kew Gardens, England, UK. March
Crocodile monitor (Varanus salvadorii) close up eye, captive, occurs in New Guinea
Pine grosbeak (Pinicola enucleator), male feeding juvenile, Finland, July
Black grouse (Lyrurus tetrix), male at lek, Finland, April
Long-tailed duck (Clangula hyemalis), males in flight, Finland, April
Kingfisher (Alcedo atthis) male, after diving, taking off from water with fish, a Common Roach (Rutilus rutilus) Lorraine, France, August
Common swift (Apus apus) group flying overhead, Bradford-on-Avon, Wiltshire, UK, May
Common European toad tadpoles (Bufo bufo), sequence showing development of limbs. Derbyshire, UK. August. Digital composite
White-throated magpie-jay (Calocitta formosa) Nicoya peninsula, Costa Rica. February
Adult Aye-aye (Daubentonia madagascariensis) active in forest canopy at night. Dry deciduous forest near Andranotsimaty. Daraina, northern Madagascar. Critically Endangered
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Canebreak treefrog (Aplastodiscus sp) Guainumbi Private Reserve, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Eurasian hoopoe (Upupa epops) at nest in stone wall with hungry chicks gaping, Vendee, France, June
Clarion Island Whip Snake (Masticophis anthonyi), IUCN Critically Endangered, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage
Grey wolf (Canis lupus), adult with cub, captive, USA
Grass snake(Natrix natrix) coiled round eggs, Alsace, France
Scarlet ibis (Eudocimus ruber), two birds perched on bushes, Coro, Venezuela. Small repro only
Scarlet ibis (Eudocimus ruber), in flight, Coro, Venezuela
Nurse shark (Ginglymostoma cirratum) portrait, resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas. The Bahamas National Shark Sanctuary
Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
Southern Viscacha (Lagidium viscacia) at rest, Isla Incahuasi in the Salar de Uyuni, altiplano, Bolivia
Zebra (Equus quagga) leaping during stampede, Serengeti, Tanzania. Vignette added and right edge expanded
Emerald toucanet (Aulacorhynchus prasinus) Rio Platano Biosphere Reserve and UNESCO World Heritage Site, La Mosquitia, Honduras
Nile crocodile (Crocodylus niloticus) Chobe National Park, Botswana, June
Galapagos sea lion (Zalophus wollebaeki) pup on shore, Galapagos
Giant panda (Ailuropoda melanoleuca) female, Huan Huan, holding baby age three months, Beauval Zoo, France, November 2017
Fin whale (Balaenoptera physalus) just below surface, south Barcelona coast, Spain, Mediterranean Sea
Whale headed / Shoebill (Balaeniceps rex) vocalising. Swamps of Mabamba, Lake Victoria, Uganda
Fin whale (Balaenoptera physalus) with ctenophore in front of its mouth, south Barcelona coast, Spain, Mediterranean Sea
Short-finned pilot whale (Globicephala macrorhynchus) showing underside whilst vertical underwater, South Tenerife, Canary Islands, Atlantic, October
Thomass Langur (Presbytis thomasi), animal endemic to northern Sumatra. These primates have relatively long arms and legs, adapted for an acrobatic life in the crowns of trees
Bairds tapir (Tapirus bairdii) Corcovado National Park, Costa Rica, May. Endangered
Stellers Sea Eagle (Haliaeetus pelagicus) with prey in beak in mid-air, Hokkaido, Japan, February
Close up if a Great hammerhead shark (Sphyrna mokarran) swimming over sandy seabed, South Bimini, Bahamas. The Bahamas National Shark Sanctuary, West Atlantic Ocean
Hoffmanns Two-toed sloth (Choloepus hoffmanni) mother and baby, aged 2 months, in tree, Costa Rica. Rescued and released by Aviarios Sloth Sanctuary
Proboscis monkey (Nasalis larvatus) mother suckling infant, Sabah, Malaysia
Harpy eagle (Harpia harpyja) close up head portrait, captive