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Snow buntings (Plectrophenax nivalis) flock in flight during a snowstorm, New York, USA, January
Snow Geese (Chen caerulescens) large flock in pool at Bosque Del Apache National Wildlife Refuge with setting full moon at sunrise in December, New Mexico, USA. December
Snowy Egret (Egretta thula) adult in breeding plumage taking flight, Viera Wetlands, Florida, USA, March
Spotted sandpiper (Actitis macularius), with head submerged as it jabs at underwater prey, Caroline, New York, USA, May
Western grebe (Aechmophorus occidentalis) adult with begging chick riding on its back, peeking out from behind adults neck, Bear River Migratory Bird Refuge, Utah, USA, May
Black-bellied whistling-ducks (Dendrocygna autumnalis), group perched amid Spanish moss on bald cypress tree, Lakeland, Florida, USA, January
Ovenbird (Seiurus aurocapillus) bringing nest material to its nest on the forest floor, Ithacs, NY, USA
Southern Sea otter (Enhydra lutris) mother holding young pup (less than one week in age), Monterey Bay, California, USA
North American River Otter (Lontra canadensis) two swimming near the surface with autumn leaves reflected in water, Acadia National Park, Maine, USA, October
Rainbow trout (Oncorhynchus mykiss) in green turbid waters, Gunnison River, Colorado, USA, April
Sea otter (Enhydra lutris) resting amongst sea ice, Alaska, USA, June
RF - American Bison (Bison bison) male in snow covered in frost, standing by dead trees. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January
Pack of wolves (Canis lupus). Yellowstone National Park, Wyoming, USA. January
Wild Salt River stallion grazing in water, living wild in Tonto National Forest, Arizona, USA. October
Rainbow scarab (Phanaeus vindex) male on white background. Florida, USA. Controlled conditions
Grand Canyon at dusk, abstract hazy landscape, Arizona, USA, July
Moose (Alces americanus) close-up grazing, Denali National Park, Alaska, USA, September
Western diamond-backed rattlesnake (Crotalus atrox) Texas, USA, April
Series of Bald eagle (Haliaeetus leucocephalus) in flight, Homer, Alaska, USA, March. Digital composite
Milky Way rising over the south wall of the Grand Canyon, Arizona, USA. November 1 2015. Composite image
Night-blooming cereus cactus (Acanthocereus tetragonus), flower bud opening at night, Texas, USA. Sequence 7 of 7. August
Sea otter (Enhydra lutris) mother and sleeping newborn pup (aged 3 days) Monterey, California, USA. *Digitally darkened scar on mothers nose
The Witch Head Nebula, IC 2118. If you reverse the image 180 degrees, you can readily see the profile of the witch, not so recognizable upside down
Polar bear (Ursus maritimus) sow with two cubs walking past a pile of Bowhead whale (Balaena mysticetus) bones, Kaktovik, Barter Island, 1002 Area, Arctic National Wildlife Refuge, North Slope
Polar bear (Ursus maritimus) spring cub sitting upright on hind legs balancing, Bernard Spit, 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Green lacewing (Abachrysa eureka) in flight, , Tuscaloosa County, Alabama, USA Controlled conditions. May
Harlequin ladybird (Harmonia axyridis) flying, Tuscaloosa County, Alabama, USA Controlled conditions. January
Purple-striped jellyfish (Chrysaora colorata) captive, USA, January
English Springer Spaniel (show type) by red berries, wild grasses, Waterford, Connecticut, USA
Red wasp (Polistes carolina) in flight Lamar County, Texas, USA Controlled conditions. August
Monarch butterfly (Danaus plexippus) nectaring on Purple Loosestrife in wet meadow, East Haddam, Connecticut, USA
Newfoundland dog, male standing in snow. Killingworth, Connecticut, USA
Black skimmer (Rynchops niger) foraging along surf line. Tierra Verde, Florida, USA
Chocolate labrador retriever, two puppiessitting in spring flowers. Haddam, Connecticut, USA. June
View up into canopy of Aspen (Populus) trees against blue sky in autumn, Grand Staircase-Escalante National Monument, Utah, USA. October
Pika, (Ochotona princeps) bringing vegetation to Hay pile, in Bridger National Forest, Wyoming, USA, July
RF - Polar Bear (Ursus maritimus), female resting, with two cubs along a barrier island outside Kaktovik, Alaska, USA. September
Northern Cardinal (Cardinalis cardinalis), adult male perched on icy branch of Possum Haw Holly (Ilex decidua) with berries, Hill Country, Texas, USA. February
Head portrait of grey almost white Andalusian mare running in snow, Berthoud, Colorado, USA. January
Wild Mustang foal resting in wildflowers, Pryor Mountains, Montana, USA. June
Head portrait of wild dun roan Mustang mare in Sand Wash Basin, Colorado, USA
American oystercatcher (Haematopus palliatus) feeding on Calico Clam, shortly after dawn, seashore, Tampa Bay, St. Petersburg, Florida, USA. April
Formation caused by erosion in sandstone, Devils Garden, Grand Staircase-Escalante National Monument, Utah, USA. April 2014
Stormy sky above the prairie grassland, Montana, USA. June 2011
Polar bear (Ursus maritimus), adult male, outside Kaktovik, Alaska, USA
RF - Bald eagle (Haliaeetus leucocephalus) calling, captive, occurs in North America
Golden Retriever pup in snow, Holland, Massachusetts, USA
Caddisfly (Hydropsychidae) in flight and attracted to mercury vapour light, Lamar County, Texas, USA, August. August