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Southern magnolia / Bull bay (Magnolia grandiflora) flower. The opening and closing of petals controls the time beetle pollinators enter and leave the flower
Texas bluebonnets (Lupinus texensis) and Indian paintbrush (Castilleja coccinea) amongst wildflowers in ditch beside byway. Rains triggered germinaton of annual seeds. Cuero, Texas, USA. April
Tulip tree (Liriodendron tulipifera) flower, pollen and nectar fluorescing in UV light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 2/2
Southern catalpa (Catalpa bignonioides) flowers. Recently opened with yellow nectar guides, purple in older pollinated flowers. Surrey, England, UK. Native to USA
Buacaille Etive Mor and River Etive in winter, Rannoch Moor, Glencoe, Scotland, UK.December
North American porcupine (Erethizon dorsatum) in snow, Vermont, USA. (Habituated rescued individual returned to the wild)
Great horned owl (Bubo virginianus) preening two chicks in nest in Saguaro (Carnegiea gigantea) cactus, Sonoran desert, Arizona, USA
American alligator (Alligator mississippiensis) emerging from water, in evening light. Myakka River State Park, Florida, USA. March
Whale shark (Rhincodon typus) close to the camera with a golden trevally fish (Gnathanodon speciosus) and outline of another whale shark behind, in the plankton-rich green waters of La Paz bay
Berkshire piglet in a basket among squashes in early autumn; Rhode Island, USA. October
Beaver pond with beaver lodge (on the left side) and trees reflected in autumn. Acadia National Park, Maine, USA. October
Moose (Alces alces) bull, crossing mountain river after sunset. Grand Teton National Park, Wyoming, USA. October 2017
Pine tree (Pinus sp) in winter, Bryce Canyon National Park, Utah, USA, January
Snowy egret (Egretta thula) shows its breeding plumage. St. Johns Management Area, Florida, USA, March
Western screech owl (Megascops kennicottii) is camouflaged as it looks from a tree cavity. Bosque del Apache National Wildlife Refuge, New Mexico, USA, January
Soaptree yucca (Yucca elata) at sunset. White Sands National Monument, New Mexico, USA. January
Sea Otter, (Enhydra lutris), young male, Elkhorn Slough, California, USA. January
Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July
American oystercatcher (Haematopus palliatus) adult feeding chick, Long Island, New York, USA. June
American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May
Common loon (Gavia immer) adult in breeding plumage, preening in the last light of day, Michigan, USA, June
Blue jay (Cyanocitta cristata) holding an acorn in its bill whilst perched on tree stump, with autumn foliage, New York, USA, October
Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions
White pelicans (Pelecanus erythrorhynchos) at Bear River Migratory Bird Refuge with an approaching storm in May, Utah, USA, May
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male perched in marsh vegetation (Phragmites) in foreground. Bear River Migratory Bird Refuge, view toward east
Indigo Bunting (Passerina cyanea) male in breeding plumage perched in Crabapple (Malus sp.) blossom in spring, Ithaca, New York, USA, May
Loggerhead shrike (Lanius ludovicianus) on an ice-covered fence after an ice storm, Anahuac National Wildlife Refuge, Texas, USA. January
Pileated Woodpecker (Dryocopus pileatus) male, at feeding excavation in tree trunk in winter, Freeville, New York, USA, January
Pileated woodpecker (Dryocopus pileatus) male excavating in search of food, New York, USA. Digital composite
Red-breasted Mergansers (Mergus serrator) male in foreground performing courtship display to female in background (left), Aurora, New York, USA, April
Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July
Common loons (Gavia immer), two adults and chick swimming on a misty lake in early morning, Michigan, USA. June
Common redpoll (Carduelis flammea), male perched in alder (Alnus rugosa) during a snowstorm in winter, New York, USA, January
Eared grebe (Podiceps nigricollis), adult in breeding plumage, swimming amidst Water crowfoot flowers, (Ranunculus aquatilis) flowers, Benton Lake National Wildlife Refuge, Montana, USA, July
Eared Grebes (Podiceps nigricollis), adult feeds a damselfly nymph to one of two chicks riding on its back, Bowdoin National Wildlife Refuge, Montana, USA, June
Great crested flycatcher (Myiarchus crinitus), female alighting at nest cavity entrance carrying pine needles as nest material, wide angle captured with remote camera to show habitat, New York, USA
Eastern screech-owl (Megascops asio) grey morph, roosting in tree cavity in winter, Lansing, New York, USA, January
Sandhill cranes (Grus canadensis) (Florida race), adult with two small chicks crossing highway, Kissimmee, Florida, USA, March
Golden-fronted woodpecker (Melanerpes aurifrons), male, Cozad Ranch, Rio Grande Valley, Texas, USA, March
Tufted puffin (Fratercula cirrhata), adult in breeding plumage flying in preparing to land, St. Paul, Pribilof Islands, Alaska, USA, July
Great Egret (Ardea alba), about to land in water, backlit, Bolsa Chica Ecological Reserve, California, USA, October
Viera Wetlands at sunrise with Great Blue Heron (Ardea herodias) in the foreground. This man-made wetland forms part of Brevard Countys water treatment system in central Florida, USA. March
Western tanager (Piranga ludoviciana), male perched on aspen branch in spring, Mono Lake Basin, California, USA, May
Western Grebe (Aechmorphus occidentalis), pair during rushing courtship display in which they run across waters surface in synchrony, near Escondido, California, USA, January
Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October
Wilsons Phalaropes (Phalaropus tricolor) flock flying over tufa formations at Mono Lake in late summer, with reflection, California, USA. July 2013
Yellow-bellied Sapsucker (Sphyrapicus varius), male drumming, long exposure, New York, USA, April