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Waterfall during monsoon, Kudremukh National Park, Western Ghats UNESCO Natural World Heritage Site, India. January 2016
Cloud forest of the Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica. Small repro only
Tetrapod tracks preserved in rock at the Joggins Fossil Cliffs UNESCO World Heritage Site, Bay of Fundy, Nova Scotia, Canada. May 2017
Tree-like lycopodiophyte (Sigillaria) preserved in situ at Joggins Fossil Cliffs UNESCO World Heritage Site along the shore, Bay of Fundy, Nova Scotia, Canada. May 2017
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
Split level view of Brown pelicans (Pelecanus occidentalis) feeding, Tortuga Bay, Santa Cruz Island, Galapagos
Brown pelican (Pelecanus occidentalis) feeding chicks at nest, Puerto Ayora / Academy Bay, Santa Cruz Island, Galapagos
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Gough Island bunting (Rowettia goughensis) gathering nesting material. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic. Endemic. Critically endangered species
Ascidian (Rhopalaea sp) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Goby (Bryaninops sp) on coral branch, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Underwater view of African tigerfish (Hydrocynus vittatus), predatory fish in the Okavango river with Water lily (Nymphaea nouchali var. caerulea) and other aquatic plants
Aerial view of the meandering Saint George branch of the Danube river, Danube Delta Biosphere Reserve UNESCO World Heritage Site, Romania, May 2014. Photographed for The Freshwater Project
Asian wild ass (Equus kiang) herd in wetlands near Madoa, Tibetan Plateau, Qinghai, China
Tibetan antelope or Chiru (Pantholops hodgsonii), Keke Xili, Changtang, Tibetan Plateau, Qinghai, China
White-rumped Snowfinch (Montifringilla taczanowskii) with chick at nest burrow, Sanjiangyuan National Nature Reserve, Qinghai Hoh Xil UNESCO World Heritage Site, Qinghai-Tibet Plateau
Hallelujah Mountains (Floating Mountains), Zhangjiajie National Forest Park UNESCO World Heritage Site, Hunan, China
Durdle Door, with incoming tide at sunset, near Lulworth, Dorset Jurassic coast, UK. June 2017
Siberian tundra reindeer (Rangifer tarandus sibiricus) Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Frozen waterfall in Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia, April
Waterfall and river, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Putorana snow sheep (Ovis nivicola borealis) with lamb, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Siberian jay (Perisoreus infaustus) Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Meltwater stream through ice, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia, June
Galapagos sea lion (Zalophus wollebaeki) hunting tuna. A group of the sea lion bulls have learnt to herd Pelagic yellowfin tuna into a small cove, trapping them
Galapagos sea lions (Zalophus wollebaeki) hunting tuna. A group of the sea lion bulls have learnt to herd Pelagic yellowfin tuna into a small cove, trapping them
Spotfin Burrfish (Chilomycterus reticulatus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Arctic fox (Vulpes lagopus) in summer moult stealing Snow goose (Chen caerulescens caerulescens) egg, with Snow goose flapping wings cand calling, Wrangel Island, Far Eastern Russia, June
Dingo (Canis lupus dingo) fighting on a beach. Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
Little island Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia
Iceberg in the sea outside the Kangia Ilulissat Icefjord. Iceberg from the Sermeg Kujalleg Glacier, UNESCO World Heritage Site, Kalaallit Nunaat, Greenland, August 2014
The Pont St-Benezet, Palais des Papes and Rhone River at dusk, Avignon, Provence, France, October 2012
Musk ox (Ovibos moschatus) two covered in snow, Wrangel Island, Far Eastern Russia, September
Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May
Female Sea lion (Zalophus wollebaeki) with pup suckling in shallow water, Floreana Island, Galapagos
Galapagos sea lion (Zalophus wollebaeki) pup on beach, Floreana Island, Galapagos, Endangered species
Clarion Burrowing Owl (Athene cunicularia rostrata), Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
RF- Sunset on Kimmeridge Bay, Isle of Purbeck, Jurassic Coast World Heritage Site, Dorset, England, UK. November 2008
Orange cup coral (Tubastrea coccinea) Malpelo Island National Park, UNESCO World Heritage Site, Colombia, East Pacific Ocean
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Longnose hawkfish (Oxycirrhites typus) hiding in coral, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Musk ox (Ovibos moschatus) two in habitat, Wrangel Island, Far Eastern Russia, October
Musk ox (Ovibos moschatus) portrait of two standing closely, Wrangel Island, Far Eastern Russia, September
Water cascading over naturally formed tufa dam. Tufa is a kind of limestone, formed from the precipitation of carbonate minerals which builds up around mosses
Mountain gorilla (Gorilla beringei beringei) female carrying baby on her back, member of Humba group, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
RF - Laurel forest / Laurisilva. Ancient evergreen hardwood trees. Madiera. (This image may be licensed either as rights managed or royalty free.)
Galapagos giant tortoise (Chelonoidis nigra) portrait, Santa Cruz Island, Galapagos Islands, Ecuador. Endangered
Galapagos sea lion (Zalophus wollebaeki) newborn pup resting head on mothers flipper on beach, Lobos Island, Galapagos. Endangered