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Galapagos Shark (Charcharhinus galapagensis), Roca Partida islet, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Galapagos giant tortoise (Geochelone nigra porteri), Cerro El Chato, Santa Cruz Island, Galapagos Islands
Great Frigatebird (Fregata minor) arriving at nest with chick, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
European Chubb (Squalius / Leuciscus cephalus) in cave, Plitvice Lakes National Park, Croatia. November
Strangler fig (Ficus sp) growing over Hoop Pine tree(Araucaria cunninghamii) with branches knotting round each other, Morans Falls Track, Green Mountains
Sumatran orangutan (Pongo abelii) female next to infant, Gunung Leuser National Park, Indonesia, UNESCO World Heritage site, November
Sumatran orangutan (Pongo abelii) female moving between lianas, Gunung Leuser National Park, UNESCO World Heritage site, November
Band of wild Rapa Nui horses / colts, walking near Easter island heads on Ahu TongarikiBand of wild Rapa Nui horses/colts, walking near Easter island heads on Ahu Tongariki, Rapa Nui National Park UNESCO World Heritage Site, Easter Island / Rapa Nui, Chile
Indian rhinoceros (Rhinoceros unicornis) portrait. Manas National Park UNESCO World Heritage Site, Assam, India
Banyan tree (Ficus macrophylla) Valley of the Shadows, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Dingo (Canis lupus dingo) on the beach at night, Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
Indian hog deer (Hyelaphus porcinus) mother and fawn, Manas National Park UNESCO World Heritage Site, Assam, India
Aldabra Giant Tortoises (Aldabrachelys gigantea) resting in a pool to keep cool, Grand Terre, Natural World Heritage Site, Aldabra
Arctic raspberry (Rubus arcticus) in flower, Padjelanta National Park, Kvikkjokk, Greater Laponia Rewilding Area, Lapland, Norrbotten, Sweden, June
Aerial view of the Rapa river delta with distant mountains, Sarek National Park, Greater Laponia Rewilding Area, Lapland, Norrbotten, Sweden, June 2013
RF- Aerial view of river with clouds reflected in water, Sjaunja Bird Protection Area, Greater Laponia Rewilding Area, Lapland, Norrbotten, Sweden, June 2013
San Cristobal giant tortoises (Chelonoidis chatamensis), fighting, Galapaguera, San Cristobal Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) in the surf, Black Beach, Floreana Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) in water, Puerto Egas, Santiago Island, Galapagos
Galapagos lava lizard (Microlophus albemarlensis) sitting on Marine iguana (Amblyrhynchus cristatus) Punta Espinosa, Fernandina Island, Galapgos
Floreana giant tortoise hybrid descendants (Chelonoidis elephantopus) feeding on flower. These are descendants of the extinct Floreana giant tortoise, but are hybridised with other species
Galapagos green turtle (Chelonia mydas agassizii) swims over a coral reef. Darwin Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Oceanic manta (Mobula birostris). Cousins Rock, San Salvador Island, Santiago Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Galapagos barnacle blenny (Acanthemblemaria castroi) looking out from its home in an old barnacle shell. Cape Marshall, Isabela Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Western Santa Cruz giant tortoise (Chelonoidis porteri) portrait, Highlands, Santa Cruz Island, Galapagos
Alcedo giant tortoises (Chelonoidis vandenburghi) group resting in water to keep cool, Alcedo Volcano, Isabela Island, Galapagos
The Cobb, Lyme Regis, Jurassic Coast, Dorset, England, UK. December 2010
Lioness (Panthera leo) sitting during misty sunrise, Okavango Delta, Botswana
Aerial view of Siberian tundra reindeer (Rangifer tarandus sibiricus) Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Frozen waterfall, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia, May 2015
Sable (Martes zibellina) Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Arctic fox (Vulpes lagopus) in summer coat calling, Wrangel Island, Far Eastern Russia, August
Musk ox (Ovibos moschatus) two in habitat, Wrangel Island, Far Eastern Russia, September
Musk ox (Ovibos moschatus) herd, Wrangel Island, Far Eastern Russia, March
Polar bear (Ursus maritimus) mother with three very young cubs, Wrangel Island, Far Eastern Russia, March
Mountain range with rainforests in morning mist, Kudremukh National Park, Western Ghats UNESCO Natural World Heritage Site, India
Nilgai (Boselaphus tragocamelus) young bulls, Keoladeo / Bharatpur National Park UNESCO Natural World Heritage Site, India
Waterfall during monsoon, Kudremukh National Park, Western Ghats UNESCO Natural World Heritage Site, India. January 2016
Cloud forest of the Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica. Small repro only
Tetrapod tracks preserved in rock at the Joggins Fossil Cliffs UNESCO World Heritage Site, Bay of Fundy, Nova Scotia, Canada. May 2017
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
Split level view of Brown pelicans (Pelecanus occidentalis) feeding, Tortuga Bay, Santa Cruz Island, Galapagos
Brown pelican (Pelecanus occidentalis) feeding chicks at nest, Puerto Ayora / Academy Bay, Santa Cruz Island, Galapagos
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Gough Island bunting (Rowettia goughensis) gathering nesting material. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic. Endemic. Critically endangered species