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Scalesia pedunculata forest, Cerro Crocker Region, Santa Cruz Island, Galapagos
Nazca booby (Sula granti) on the coast of Wolf Island, Galapagos
Magnificent frigatebirds (Fregata magnificens) diving for fish prey, Borrero Bay, Santa Cruz Island, Galapagos
Magnificent frigatebird (Fregata magnificens) attacking Blue footed booby (Sula nebouxii) feeding chick. This bird is trying to steal the food from the parent. Santa Cruz Island, Galapagos
Galapagos hawk (Buteo galapagoensis) flying, carrying an iguana carcass, Punta Espinosa, Fernandina Island, Galapagos
Galapagos dove (Zenadia galapagensis) on head of Galapagos giant tortoise, (Chelonoidis darwini) Santiago Highlands, Galapagos
Feral domestic cattle, Puerto Fragata, Isabela Island, Galapagos
Feral kitten with sleeping Galapagos sea lion (Zalophus wollebaeki) Puerto Fragata, Isabela Island, Galapagos
Espanola saddelback tortoise (Chelonoidis hoodensis) pair mating, Espanola Island, Galapagos
Vegetarian finch (Platyspiza crassirostris) snips petals from a hardy vine, Fernandina Island, Galapagos
Short-eared owl (Asio flammeus) in flight, Santa Fe Island, Galapagos
Galapagos penguin (Spheniscus mendiculus) and Marine iguana (Amblyrhynchus cristatus) male, Isla Tortuga, Isabela Island, Galapagos
Panamic Cushion Sea Star (Pentaceraster cumingi) and a school of Yellow-tailed Grunt (Anisotremus interruptus), Galapagos Island, Ecuador. December
Magnificent frigatebird (Fregata magnificens) in flight over sea between Isabela Island and Fernandina Island, Galapagos
Galapagos sea lion (Zalophus wollebaeki) jumping out of Pacific ocean. Sombrero Chino, Santiago Island, Galapagos
Orca (Orcinus orca) pod hunting Sealion, tossing it out of the sea, Punta Norte, Valdez Peninsula, Argentina. April
Orca (Orcinus orca) hunting South American sealion (Otaria flavescens) close to shore. Punta Norte, Valdez Peninsula, Argentina. April
Bamboo growing beside narrow mountain track. Gaoligongshan National Nature Reserve, Three Parallel Rivers of Yunnan Protected Areas, Yunnan Province, China. May 2009. Digitally stitched panorama
Scalloped hammerhead shark (Sphyrna lewini) swims through a school of Pacific creolefish (Paranthias colonus). Wolf Island, Galapagos National Park, Galapagos Islands.. East Pacific Ocean
Marine Iguanas (Amblyrhynchus cristatus) basking. Fernandina Island, Galapagos Islands
Forest in autumn colours, Plitvice Lakes National Park, UNESCO World Heritage Site, Central Croatia. Croatia
Red-footed booby (Sula sula) perched in tree. Genovesa Island, Galapagos
Nazca booby (Sula granti), pair on rocky shore at dawn. Punta Suarez, Espanola Island, Galapagos. May 2015
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Marine iguana (Amblyrhynchus cristatus), Punta Vicente Roca, Isabela Island, Galapagos
Sooty albatross (Phoebetria fusca) nesting amongst Blechnum palmiforme tree ferns. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display sequence. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Yellow warbler (Setophaga petechia aureola) on lava rocks. Isabela Island, Galapagos. May
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Rainforest and river outside the Batak village of Sitio Manggapin in Cleopatra sRainforest and river outside the Batak village of Sitio Manggapin in Cleopatras Needle Critical Habitat, Palawan, Philippines
Mangroves on Tupeti Island, Southern Lagoon, Forgotten Coast, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Coral formations at low tide in Prony Bay in the Southern Lagoon, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Rock flagtail (Kuhlia rupestris) fish in river on the Southern Lagoon, Forgotten Coast, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Galapagos green turtle (Chelonia mydas agassizi) swimming in coastal waters of San Cristobal Island, Galapagos
Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Flightless cormorant (Phalacrocorax harrisi), adult and two chicks. Beagle Crater, Isabela Island, Galapagos
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Mountain landscape reflected in lake, Basongcuo National Park, Qinghai-Tibet Plateau, Tibet, November 2011
Suspension bridge across the river, entrance to Gunung Mulu National Park UNESCO Natural World Heritage Site, Malaysian Borneo
Landscape modified by the erosion process. Goreme National Park and the Rock Sites of Cappadocia UNESCO World Heritage Site. Turkey. December 2006
Silver Drummer (Kyphosus sydneyanus) fish off the coast of Neds Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
River through Virgin Komi Forests UNESCO World Heritage site, the largest virgin forests in Europe. Ural Mountains, Komi Republic, Russia. August 2016
Freshwater isopods (Acanthogammarus victorii) feeding on seabed, Lake Baikal, Siberia, Russia
Young Stone sculpin (Paracottus knerii), on endemic sponge (Lubomirskia baicalensis), Lake Baikal, Siberia, Russia
Clarion Lizard (Urosaurus clarionensis) displaying, IUCN Vulnerable, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Tropical Flounder (Bothus mancus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean